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ACCESSIONPLANT NAMETAXONOMYORIGINGENEBANKIMAGEAVAILABILITYRECEIVEDSOURCE TYPESOURCE DATECOLLECTION SITECOORDINATESELEVATIONHABITATIMPROVEMENT LEVELNARRATIVE
0PI 330517IntermediumHordeum vulgare L. subsp. vulgare Reading, United KingdomNSGCKERNELS1968DONATED05/16/196851.45000000, -0.96666700Uncertain improvement status1246010PI 330517
1PI 184882IntermediumHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1949DEVELOPED58.98333300, 13.50000000Genetic material1159797PI 184882
2GSHO 677intermediumHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1995DONATED02/01/1995Genetic materialH. Intermedium1512153GSHO 677
3GSHO 1764int-b.3Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialThe spike appears similar to the six-rowed spike, but developmental irregularities occur commonly in the lower portion of the spike. All lateral spikelets are reduced in size, and the lemma awn is short or reduced to a pointed tip. Commonly, only lateral spikelets in the middle of the spike set seed. Mutant plants have reduced vigor and tillering. Inheritance: Monofactorial recessive. Located in chromosome 5HL [7L], near the raw1 (smooth awn 1) locus based on linkage drag. Previous nomenclature and gene symbolization: None. Mutational events: int-b.3, -b.6 in Bonus; int-b.75 in Bonus. Mutant used for description and seed stocks: int-b.3 in Bonus (GSHO 1764); int-b.3 in Bowman (PI 483237)*7 (GSHO 2129).1006044GSHO 1764
4GSHO 1767int-f.19Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialThe spike appears to be six-rowed, but the lateral spikelets are much smaller, less than half the size of the central spikelets. Lateral spikelets are pointed and often have short awns. Seed set occurs in the lateral spikelets in the upper third of the spike. The base of the spike has shortened rachis internodes and appears erectoides-like. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: int-f.19 in Foma. Mutant used for description and seed stocks: int-f.19 in Foma (GSHO 1767); int-f.19 in Bowman (PI 483237)*3 (GSHO 2270).1006080GSHO 1767
5GSHO 1768int-h.42Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialLateral spikelets are enlarged and have an inconspicuously pointed apex, but they do not set seed. Induced mutants show early heading and have an elongated basal internode. The spike appears lax with shortened rachis internodes at the base. The Bowman backcross-derived line is slightly shorter (5/6 normal) and produces extra spikelets (up to five fertile ones) at several rachis nodes in the lower half of the spike. Fusion of some spikelets results in double and occasionally triple kernels. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: int-h.42, -h.43, -h.44 in Kristina; int-h.83 in Bonus (PI 189763). Mutant used for description and seed stocks: int-h.42 in Kristina (GSHO 1768); int-h.42 in Bowman (PI 483237)*5 (GSHO 2271).1006081GSHO 1768
6GSHO 1769int-i.39Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialLateral spikelets are enlarged and slightly pointed at the apex, but they fail to set seed. The tip of the spike may have very short rachis internodes and appears very dense or fasciated. The Bowman backcross-derived line exhitbits reduced plant height (5/6 normal), reduced spike length, and a distinct elongation of the basal rachis internode. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: int-i.39 in Kristina. Mutant used for description and seed stocks: int-i.39 in Kristina (GSHO 1769); int-i.39 in Bowman (PI 483237)*4 (GSHO 2272).1006102GSHO 1769
7GSHO 1772int-mHordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1996DEVELOPED58.98333300, 13.50000000Genetic materialThe spike is very short and has irregular rachis internode lengths. Lateral spikelets are enlarged and pointed, but they do not set seed. Spikelet density at the base of the spike is increased. Rachis internodes at the tip of the spike are very short, and the spike appears to have two or three fused or fasciated terminal spikelets. Tillering of int-m plants is increased and heading is slightly earlier. Inheritance: Monofactorial recessive. Location is unknown. Previous nomenclature and gene symbolization: None. Mutational events: int-m.85 in Bonus; int-m.la (GSHO 1772) in Lamont (PI 512036). Mutant used for description and seed stocks: int-m.85 in Bonus (GSHO 1771); int-m.85 in Bowman (PI 483237)*5 (GSHO 2273); int-m.la in Bowman (PI 483237)*4 (GSHO 2274).1006123GSHO 1772
8GSHO 2492sls1.aHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1996DONATED04/1996Genetic materialSterile lateral spikelets become progressively smaller toward the tip of two-rowed spikes in plants, which express the Vrs1.b and int-c.b alleles at the six-rowed spike 1 and intermedium spike-c loci, respectively. The terminal rachis nodes generally have rudimentary lateral spikelets. Expression of the sls is affected by genetic background, but often is stronger in late tillers. The trait can be observed only in some two-rowed segregates from crosses between six- and two-rowed cultivars. The sls phenotype can be observed in plants of the Bowman backcross-derived line for the msg1.ca gene. Two-rowed segregates in the Harrington/Morex doubled haploid population show the typical range of expressions associated various combinations of sls and int-c alleles.1545056GSHO 2492
9GSHO 1987int-e.58Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DEVELOPEDGenetic materialPheno: Intermedium spike-e; Gener: BC5F3; Cross: I94-462; Source: 95FGH874 Bowman backcross. 1534475GSHO 1987
10GSHO 2003int-c.5Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DEVELOPEDGenetic materialPheno: Intermedium spike-c; Gener: BC5F3; Cross: I94-480; Source: 95FGH889 Bowman backcross. 1534400GSHO 2003
11GSHO 2055int-a.1Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DEVELOPEDGenetic materialPheno: Intermedium spike-a; Gener: BC4F3; Cross: I92-592; Source: 93FGH716 Bowman backcross. 1534220GSHO 2055
12GSHO 2129int-b.3Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DEVELOPEDGenetic materialPheno: Intrmedium spike-b; Gener: BC4F3; Cross: I93-633-2; Source: 95FGH986 Bowman backcross. 1534513GSHO 2129
13GSHO 2270int-f.19Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DEVELOPEDGenetic materialPheno: Intermedium spike-f; Gener: BC2F4; Cross: I91-602; Source: 93FGH920 Bowman backcross. 1534441GSHO 2270
14GSHO 2271int-h.42Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DEVELOPEDGenetic materialPheno: Intermedium spike-h; Gener: BC4F3; Cross: I95-549; Source: 95FGH192 Bowman backcross. 1534575GSHO 2271
15GSHO 2272int-i.39Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DEVELOPEDGenetic materialPheno: Intermedium spike-i; Gener: BC3F3; Cross: I93-750; Source: 94LAG1091-1 Bowman backcross. 1534442GSHO 2272
16GSHO 2273int-m.85Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DEVELOPEDGenetic materialPheno: Intermedium spike-m; Gener: BC4F3; Cross: I93-751; Source: 94FGH665 Bowman backcross. 1534178GSHO 2273
17GSHO 2274int-m.1aHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DEVELOPEDGenetic materialPheno: Intermedium spike-m; Gener: BC3F4; Cross: I90-363; Source: 93FGH837 Bowman backcross. 1534250GSHO 2274
18GSHO 1421dwarf, awnless, intermediumHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1995DONATED02/01/1995Genetic materialLk Lk2, dwarf, awnless, intermedium1514487GSHO 1421
19GSHO 170intermedium a.1Hordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1995DONATED01/01/1995Genetic material1510725GSHO 170
20GSHO 343buff seed, no appendages on lateralsHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1995DONATED02/01/1995Genetic materialmix elevated hood and awn type, n v b k R S Re Re re2 re2 The inheritance of color on the cross Hordeum Intermedium cornutum X Hordeum distichon Nudum, H.S. Smith M.S. Thesis Colorado State Univ.1512168GSHO 343
21GSHO 72high fertility intermediumHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1995DONATED01/01/1995Genetic materialHigh fertility intermedium, BB, NN, kk, RR, ss, purple plant color lhlh, VV.1510667GSHO 72
22GSHO 1763int-a.1Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DONATED05/1995Genetic material1545188GSHO 1763
23GSHO 1765int-c.5Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DONATED05/1995Genetic material1545295GSHO 1765
24GSHO 1766int-e.58Hordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DONATED05/1995Genetic material1545296GSHO 1766
25GSHO 1773int-m.1aHordeum vulgare L. subsp. vulgare North Dakota, United StatesNSGCKERNELS1995DONATED05/1995Genetic material1545200GSHO 1773
26GSHO 153vrs1.cHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialThe spike has long awns on the lemma of the central spikelet and awn-like appendages on the lemma of the lateral spikelets. Lateral spikelets have perfect flowers and complete fertility. The lr trait is expressed only in six-rowed barley, and heterozygotes have a short awn on the lemma of lateral spikelets. Since recombination with neither vrs1.a(v) nor Vrs1.b (V) alleles was observed, the lr or vrs1.c phenotype is likely controlled by an allele at the vrs1 locus. The Bowman backcross- derived line for the vrs1.c gene has short, stiff awns on the central spikelets (about 1/3 normal length) and no awns on the lateral spikelets.Inheritance: An incomplete dominant in the multiple allelic series at the vrs1 (six-rowed spike 1) locus. Located in chromosome 2HL, about 15.4 cM distal from the Gth (toothed lemma) locus, and 26.5 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Reduced lateral spikelet appendage on the lemma=lr. Reduced lateral spikelet appendage on the lemma=vlr. Mutational events: vrs1.c or lr in Hordeum intermedium var Nudihaxtoni. Mutant used for description and seed stocks: vrs1.c or lr in Hordeum intermedium var Nudihaxtoni (PI 32368, GSHO 153); vrs1.c in Bowman (PI 483237)*7(GSHO 1908).1510717GSHO 153
27GSHO 196vrs1.aHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialAlleles at this complex locus modify development of the lateral spikelets and the associated lemma awn. The vrs1.a allele (v gene) is present in most six-rowed cultivars and produces well-developed lateral spikelets. The lemma awn of lateral spikelets will vary from 3/4 to nearly as long as those of central spikelets, depending upon alleles present at other loci. The Vrs1.b allele (V gene, distichon) is present in many two-rowed cultivars and reduces lateral spikelets to sterile bracts with a rounded tip. The Vrs1.t allele (Vt gene, deficiens) causes an extreme reduction in the size of lateral spikelets. The lr or vlr (vrs1.c) gene in Nudihaxtoni and Bozu types will not recombine with the vrs1.a allele and produces phenotypes similar to the Vrs1.d allele (Vd gene) of Svanhals. The series of induced mutants in two-rowed barley called hex-v and Int-d mutants differ in the size of lateral spikelets, but they interact with the vrs1.a allele as incomplete dominants. Many heterozygous combinations with vrs1.a have a pointed tip on the lemma of sterile lateral spikelets. Alleles at the int-c (intermedium spike-c) locus modify lateral size in the presence of vrs1.a, Vrs1.b, and Vrs1.d, but not when Vrs1.t is present. Inheritance: A multiple allelic series, incomplete dominant allele interactions based on the size and shape of lateral spikelets. Located in chromosome 2HL, about 30.5 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization:Two-row vs six-row=Zz. Six-row vs two-row=Aa. Two-row=D. Six-row vs two-row=Vv. Six-row vs two-row (distichon) vs two-row (deficiens)=A,as,af. Redcd lateral spikelet append. on the lemma=lr. Allelic series v,Vd,V,Vt. Hex. mut. Intermed. spike-d=Int-d. Reduced lat. spike. append. on lemma-vlr. Mut.events:vrs1.a ... vrs1.o(v1b) in New Golden.Mut. used for descr.:vrs1.a...(GSHO 1907).1510659GSHO 196
28GSHO 346Vrs1.dHordeum vulgare L. subsp. vulgare Colorado, United StatesNSGCKERNELS1994DONATED05/1994Genetic materialPlants homozygous for Vrs1.d have more lateral kernels than those with Vrs1.b (V), which is the common allele in two-rowed barley. Homozygosity for the Vrs1.d allele and the int-c.b (formally i) allele at the int-c (intermedium-c) locus prevents the development of ovules and awns in lateral spikelets. The Vrs1.d allele interacts with the Int-c.a (formally l) allele to cause development of a few fertile lateral spikelets. Presence of the Int-c.h (formally lh) allele contributes to a greater number of lateral kernels than Int-c.a. Inheritance: A partially dominant allele in the multiple allelic series at the vrs1 (six-rowed spike 1) locus. Located in chromosome 2HL, about 15.4 cM proximal from the Gth (toothed lemma locus, and 30.5 cM f from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Two-rowed barley of Hordeum distichon=Vd. Renamed Vrs1.d based on allelism tests. Mutational events: Vrs1.d or Vd in Svanhals (PI 5474). Mutant used for description and seed stock: Vrs1.d or Vd in Svanhals (GSHO 346).1512104GSHO 346
29GSHO 478ert-f.18Hordeum vulgare L. subsp. vulgare Skåne län, SwedenNSGCKERNELS1994DEVELOPED58.98333300, 13.50000000Genetic materialSpikes are semicompact with average rachis internode length measurements from 2.2 to 2.6 mm. Culm length is about 5/6 of normal. Sterile lateral spikelets are larger than those of normal sibs, but apices are rounded. Inheritance: Monofactorial recessive, but some partial dominance is expressed. Located in chromosome 4HS?, an association with the int-c (intermedium spike-c) locus is reported, but the association could be caused by a pleiotropic effect of the ert-f gene and not linkage. Previous nomenclature and gene symbolization: Erectoides-18=ert-18. Mutational events: ert-f.18 in Bonus; ert-f.86, -f.145 in Bonus. Mutant used for description and seed stocks: ert-f.18 in Bonus (GSHO 478); ert-f.18 in Bowman (PI 483237)*6 (GSHO 2004).1512140GSHO 478
30GSHO 684Vrs1.tHordeum vulgare L. subsp. vulgare Utah, United StatesNSGCKERNELS1994DEVELOPEDGenetic materialPlants homozygous for Vrs1.t (Vt) allele always have a deficiens phenotype regardless of the allele present at the int-c (intermedium-c) locus in chromosome 4. Deficiens is characterized by the extreme reduction in size of the lateral spikelet. The Vrs1.t allele in Engleawnless does not recombine with the Lks1.a allele at the Lks1 (awnless) locus because a short paracentric inversion is present in this stock. Inheritance: The Vrs1.t allele is the most dominant allele in the multiple allelic series at the vrs1 (six-rowed spike 1) locus, but interacts as an incomplete dominant with some alleles. Located in chromosome 2HL, about 15.4 cM distal from the Gth (toothed lemma) locus, and 30.5 cM distal from the eog (elongated outer glume) locus. Previous nomenclature and gene symbolization: Hordeum deficiens=Vt. Renamed Vrs1.t based on allelism tests. Mutational events: Vrs1.t or Vt in White Deficiens (CIho 7316). Mutant used for description and seed stocks: Vrs1.t or Vt in White Deficiens (GHSO 684); Vrs1.t in Bowman (PI 483237)*8(GSHO 1909).1512235GSHO 684
31GSHO 774vrs3.fHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialThe upper half of the spike appears to be six-rowed, but the lateral spikelets in the lower half are reduced in size with the basal portion being two-rowed. Awns on the lateral spikelets range from normal near the top, to awnletted or pointed in the middle, to awnless at the base of the spike. Lateral spikelets set seed only in the upper two-thirds of the spike. Lateral spikelets may be pedicellated in some stocks. Central spikelets often have double awn, one on the lemma and one on the palea. The rachilla may be deformed. Inheritance: Monofactorial recessive. Located in chromosome 1HL [5L]; near the centromere and proximal from the ert-b (erectoide-b) locus. Previous nomenclature and gene symbolization: Six-rowed spike 3=v3. Intermedium spike-a=int-1. Intermedium spike-g=int-g. Mutational events: vrs3.f in Hakata 2 (Kmut 213, OUM292), vrs3.g (Xb 388.8, Piro-1), vrs3.h (Xd 160.46, Piro-3), vrs3.i (X15 1463, Piro-5), vrs3.j (X17 3555, Piro-8) in Piroline (PI 539132); int-a.1 in Bonus (PI 189763); int-a.2 in Bonus; int-a.8, -a.9, -a.10, -a.64 in Bonus, -a.14, -a.17, -a.21, -a.27, -a.30, -a.31, -a.32, -a.34, -a.35, -a.37 in Foma (CIho 11333), -a.46, -a.51, -a.52, -a.54, -a.55, -a.59, -a.61 in Kristina; int-a.71, -a.74, -a.77, -a.79, -a.86, -a.88 in Bonus, -a.102, -a.103 in Hege. Mutant used for description and seed stocks: vrs3.f in Hakata 2 (GSHO 774); int-a.1 in Bonus (GSHO 170); vrs3.f in Bowman (PI 483237)*7 (GSHO 2056); int-a.1 in Bowman*5 (GSHO 2055).1513988GSHO 774
32GSHO 833lnt1.aHordeum vulgare L. subsp. vulgare JapanNSGCKERNELS1994DEVELOPEDGenetic materialThe tiller number is reduced to 2 to 4 per plant. These tillers are formed soon after seedling emergence; hence, no late-emerging tillers are observed. Culms are thick and stiff and leaves are dark green. Spike malformations occur in most environments. The spike may have irregular rachis internode lengths and is relatively short. The lower portion of the spike appears more dense. Lateral spikelets in two-rowed cultivars are enlarged and have a pointed apex. Plants homozygous for a recessive allele at the lnt1 locus head slightly earlier than normal sibs. Inheritance: Monofactorial recessive. Located in chromosome 3HL, about 30.5 cM distal from the uzu (uzu) locus. Previous nomenclature and gene symbolization: Reduced number of tillers=rnt. Intermedium spike-l=int-l. Mutational events: lnt1.a in Mitake (OUJ408); int-l.81 in Bonus (PI 189763). Mutant used for description and seed stocks: lnt1.a in Mitake (GSHO 833); int-l.81 in Bonus (GSHO 1771); lnt1.a in Bowman (PI 483237) *8 (GSHO 1984); int-l.81 in Bowman*6 (GSHO 1961).1514004GSHO 833
33CIho 7137Intermedium VVIhIhHordeum vulgare L. subsp. vulgare Utah, United StatesNSGCKERNELS1944DEVELOPEDGenetic material1038816CIho 7137
34CIho 7138Intermedium VVIhIhHordeum vulgare L. subsp. vulgare Utah, United StatesNSGCKERNELS1944DEVELOPEDGenetic material1038820CIho 7138
35CIho 6029CI 6029Hordeum vulgare L. subsp. vulgare Idaho, United StatesNSGCKERNELS1933DEVELOPEDBreeding materialIntermedium.1034642CIho 6029
36CIho 6116Orel IntermediumHordeum vulgare L. subsp. vulgare Idaho, United StatesNSGCKERNELS1933COLLECTED45.00000000, -115.00000000Breeding material1035148CIho 6116
37CIho 3187Regressive 2Hordeum vulgare L. subsp. vulgare Virginia, United StatesNSGCKERNELS1923DEVELOPEDGenetic materialSegregate which will not throw intermedium when crossed with distichon.1023567CIho 3187
38CIho 3188Regressive 38Hordeum vulgare L. subsp. vulgare Virginia, United StatesNSGCKERNELS1923DEVELOPEDGenetic materialSegregates do not yield intermedium when crossed with distichon.1023570CIho 3188
39CIho 2210MortoniHordeum vulgare L. subsp. vulgare Minnesota, United StatesNSGCKERNELS1921DEVELOPEDBreeding material1018905CIho 2210
40CIho 2214CI 2214Hordeum vulgare L. subsp. vulgare Minnesota, United StatesNSGCKERNELS1920DEVELOPEDBreeding material1018933CIho 2214
41PI 38357496Thinopyrum intermedium (Host) Barkworth & D. R. Dewey subsp. intermedium TurkeyW61972COLLECTEDPulumur. Basalt strewn, gravel soil on steep slope east side Wild materialGrowing in association with Phleum, Festuca ovina, and, Hordeum bulbosum 10 1287781PI 383574
42W6 57729ID340-5Plantago patagonica Jacq. Idaho, United StatesW62020Wild materialSOS 92nd Shipment2105027W6 57729