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ACCESSIONPLANT NAMETAXONOMYORIGINGENEBANKIMAGEAVAILABILITYRECEIVEDSOURCE TYPESOURCE DATECOLLECTION SITECOORDINATESELEVATIONHABITATIMPROVEMENT LEVELNARRATIVE
0PI 691715BJP/IM/2014/003Ilex mucronata (L.) M. Powell et al. Maine, United StatesNA2014COLLECTED08/13/2014Edge of woods on south side of Hwy. 189, across the road from a cemetery at the intersection of 189 with Timber Cove Road. Approximately 13 km southwest of Lubec.44.80087000, -67.12674000Edge of woods on south side of highway.Wild material1923674PI 691715
1PI 691727WLP 1261Ilex verticillata (L.) A. Gray New York, United StatesNAFRUIT2011COLLECTED09/12/2011Hanging Bog Game Management Area, just west of Hanging Bog, Allegany County.42.30500000, -78.25600000632Marsh. Associated with Aronia, Aster, Cornus amomum, Eutrochium, Salix, Solidago, Spiraea alba, Viburnum cassinoides, Viburnum dentatum, and Viburnum lentago.Wild material1891481PI 691727
2PI 691734WLP 1262Ilex opaca Aiton North Carolina, United StatesNA2008COLLECTED11/05/200821061922752PI 691734
3PI 296109Ilex fargesii Franch. United Kingdom Historic1990COLLECTED1681379PI 296109
4PI 287736'Golden Gem'Ilex crenata Thunb. British Columbia, Canada Historic1987COLLECTED1689796PI 287736
5PI 479432NA 51056Ilex crenata Thunb. JapanNANot Available1982COLLECTED09/15/198244.88333333, 141.91666667Wild materialSnow cover 1-2m per annum. Minimum temperature -17C. Shrub evergreen, 1.2m. Leaves lustrous above. Fruit black. Seed from one plant moved from wild.1374368PI 479432
6PI 479433JH-310-82Ilex crenata Thunb. var. paludosa (Nakai) H. Hara Japan Historic1982COLLECTED10/08/1982Wild materialSnow 2-3m, 58 days of snow cover 50cm or deeper. Minimum temperature -20C. Cuttings collected from one male plant.1374369PI 479433
7PI 479434JH-305-82Ilex crenata Thunb. var. paludosa (Nakai) H. Hara Japan Historic1982COLLECTED10/08/198240.55000000, 140.96666667Wild materialSnow 2-3m; 58 days covered by 50cm or deeper. Minimum temperature -20C. Shrub evergreen, 1.2-1.7m tall. Branches many, low spreading. Bark smooth, gray. Leaves lustrous above, pale beneath. Fruit nearly black. Growing in marsh on edge of small lake. Seed from several plants.1374370PI 479434
8PI 479435JH-267-82Ilex leucoclada (Maxim.) Makino Japan Historic1982COLLECTED10/05/198241.81666667, 140.80000000Cultivated materialSnow cover for 2 days by snow 50cm or deeper. Minimum temperature -18C. Cuttings collected from a female plant moved from Kikonai-machi where it was native.1374371PI 479435
9PI 479436JH-312-82Ilex leucoclada (Maxim.) Makino Japan Historic1982COLLECTED10/08/198240.55000000, 140.96666667Wild materialSnow cover 2-3m; 58 days covered by 50cm or deeper. Minimum temperature -20C. Cuttings collected from a male plant.1374372PI 479436
10PI 479437JH-326-82Ilex leucoclada (Maxim.) Makino Japan Historic1982COLLECTED10/09/198240.65000000, 140.83333333Wild materialSnow 2-3m; 58 days covered by snow 50cm or deeper. Minimum temperature -20C. Growing in wooded area. Cuttings from 2 male plants.1374373PI 479437
11PI 479438JH-327-82Ilex leucoclada (Maxim.) Makino Japan Historic1982COLLECTED10/09/198240.65000000, 140.83333333Wild materialSnow 2-3m; 58 days covered by snow 50cm or deeper. Minimum temperature -20C. Shrub evergreen, 60-90cm tall, erect habit, stems green. Leaves coriaceous deep green above, pale beneath. Fruit dark red, globose. Growing in Fagus crenata woodland. Cuttings from two female plants.1374374PI 479438
12PI 479439JH-390-82Ilex macropoda Miq. Japan Historic1982COLLECTED10/15/1982Foot of Mt Nyokoo, near Zyakko Falls, Nikko, Tochigi-ken, HonshuWild materialTree deciduous, 7.5m tall. Branches wide spreading. Bark smooth, whitish to gray. Leaves dark green above with impressed veins, pale beneath and lustrous. Fruit purplish red, sweet to taste. Growing in woodland on dry ridge.1374375PI 479439
13PI 479440JH-20-82Ilex rugosa F. Schmidt Japan Historic1982COLLECTED09/14/198245.08333333, 141.71666667Wild materialShrub evergreen, decumbent, 31-38.7cm. Branches rooting along, greenish. Leaves dark green aboe, rugose, light green green beneath. Fruits orange-red. Growing with Viburnum furcatum, Abies sacchalinensis, Magnolia, Quercus mongolica, Acanthopanax, Kalopanax. Seed from several plants.1374376PI 479440
14PI 479441JH-82-82Ilex rugosa F. Schmidt Japan Historic1982COLLECTED09/18/198244.05000000, 145.00000000Wild materialSnow fall 2m per annum. Growing with trees similar to JH-75. Seed collected from several plants.1374377PI 479441
15PI 479442JH-112-82Ilex rugosa F. Schmidt Japan Historic1982COLLECTED09/21/198243.30000000, 145.60000000Wild materialSnow cover 50cm or deeper for 30 days. Minimum temperature -23C. Shrub evergreen decumbent, 25-30cm tall. Stems declined. Twigs green. Leaves deep green above, lighter beneath. Fruit bright red. Growing in mixed woodland. Seed from several plants.1374378PI 479442
16PI 479443JH-102-82Ilex sugerokii Maxim. Japan Historic1982COLLECTED09/19/198244.05000000, 145.16666667Wild materialSnow cover 1.5m per annum. Minimum temperature -20C. Shrub evergreen, 1-1.8m tall, multistemmed from base. Branches dense. Leaves subcoriaceous, lustrous above, light green beneath. Fruit red, lustrous. Growing in open woodland in sunny situation. Seed from several plants.1374379PI 479443
17PI 479444JH-347-82Ilex sugerokii Maxim. Japan Historic1982COLLECTED10/10/198240.61666667, 140.93333333Wild materialSnow cover 2-3m, 58 days covered by 50cm or deeper. Minimum temperature -20C. Seed collected from several plants.1374380PI 479444
18PI 479445JH-348-82Ilex sugerokii Maxim. Japan Historic1982COLLECTED10/10/198240.61666667, 140.93333333Wild materialSnow cover 2-3m, 58 days covered by snow 50 cm or deeper. Minimum temperature -20c. Cuttings taken from male plants.1374381PI 479445
19PI 479446JH-349-82Ilex sugerokii Maxim. Japan Historic1982COLLECTED10/10/198240.61666667, 140.93333333Wild materialSnow cover 2-3m, 58 days covered by snow 50cm or deeper. Minimum temperature -20C. Cuttings taken from female plant.1374382PI 479446
20PI 465978 NA 28309'Sunny Foster'Ilex ×attenuata Ashe United StatesNANot Available1981COLLECTED1964Discovered on grounds of National Arboretum in 1964 by W.F. KosarCultivarTree evergreen with pale yellow variegated leaves. Similar in leaf shape, size, habit, hardiness, fruiting, and ease of culture to its parent clone 'Foster #2' except for slightly slower growth rate and the genetically stable variegated-leaf trait.1360914PI 465978 NA 28309
21PI 452266 NA 31352'Twiggy'Ilex crenata Thunb. District of Columbia, United StatesNANot Available1981DONATED01/1981CultivarFrom United States. United States National Arboretum, U.S. Department of Agriculture, Washington, D.C. Received January 1981. 'Twiggy'. Female seedling selected in 1964 by W.F. Kosar. Plants compact, globe-shaped, densely branched. Shearing not required. Growth rate slow to moderate. Leaves small, elliptic, dark green. Perennial woody. Cultivated. Plants. [For a time, this was not considered to be I. crenata, but a hybrid of I. crenata and I. yunnanensis, per information available from WFK; however, information regarding its verification as a cultivar of I. crenata was found buried in the Names table record, although it does not contain a reference to a published work. It may be the result of testing by Frank S. Santamour, Jr. Due to its distinctiveness it was thought for a while to be a genuine interspecific hybrid. Through later comparison of leaf flavonoids using chromatographic techniques with the reputed parent species, it was verified that 'Twiggy' was I. crenata and not a hybrid with I. yunnanensis. -SBL]1347202PI 452266 NA 31352
22PI 443301Ilex verticillata (L.) A. Gray United States Historic1980COLLECTED10/1978Carbon Co., PennsylvaniaCollected by C. Heiney.1338237PI 443301
23PI 443302Ilex verticillata (L.) A. Gray United States Historic1980COLLECTED10/1978Crawford Co., PennsylvaniaCollected by K. Hinman.1338238PI 443302
24PI 441093Ilex decidua Walter United States Historic1980COLLECTED12/1977Bell Co., TexasCollected by H. W. Everett.1336029PI 441093
25PI 399369Ilex aquifolium L. Former Serbia and Montenegro Historic1975COLLECTED1300652PI 399369
26PI 399370Ilex aquifolium L. Former Serbia and Montenegro Historic1975COLLECTED1300653PI 399370
27PI 377680 NA 35279'Jackson'Ilex verticillata (L.) A. Gray NANot Available1973CultivarFrom United States. Plants selected by O.M. Neal, Division of Plant Sciences, University of West Virginia, Morgantown, West Virginia. Received December 18, 1972. 'Jackson'. Origin same as P.I. 377678 ['Cacapon': Selected from several hundred mixed seedlings at Horticulture Farm]. Male plant, dark green foliage, produces abundant pollen. Ref: Plant Distribution List CR Form 7A. Comment: Selected from mixed seedlings from seed germination experiments.2158608PI 377680 NA 35279
28PI 377678CacaponIlex verticillata (L.) A. Gray West Virginia, United States Historic1972DONATED12/18/19721282955PI 377678
29PI 331202 NA 31489'Albert Close'Ilex hybr.NANot Available1969CultivarFrom the United States. Selection made by J.L. Creech and W.L. Ackerman at the United States Plant Introduction Station, Glenn Dale, MD. Numbered June 25, 1968. Selection from an F1 progeny resulting from a cross of Ilex cornuta, PI65860 x I. ciliospinosa, PI78144 made in 1952. 'Albert Close'. Female evergreen shrub, compact, short, pyramidal, very short internodes, dense twiggy growth. Leaves deep green, semi-glossy, spinose, ovate, 1 1/2 to 2 1/2 cm long, 1 to 1 1/2 cm wide, most 3 spines. Fruits red, round, pea-size, pedicels short.2107845PI 331202 NA 31489
30PI 331203 NA 31493'William Cowgill'Ilex hybr.Maryland, United StatesNANot Available1969DONATED06/25/1968CultivarFrom the United States. Selections made by J.L. Creech and W.L. Ackerman at the United States Plant Introduction Station, Glenn Dale, MD. Numbered June 25, 1968. Selection from an F1 progeny resulting from a cross of Ilex cornuta, PI65860 x I. ciliospinosa, PI78144 made in 1952. 'William Cowgill'. Female evergreen shrub, compact, pyramidal. Leaves medium green, glossy, spinose, lanceolate, 2 1/2 to 4 cm long, 1 1/4 to 2 cm wide, mostly 7 spines. New growth pinkish, turning green. Fruits red, round, pea-size, pedicels short, borne in clusters.1246457PI 331203 NA 31493
31PI 331204 NA 31492'Howard Dorsett'Ilex hybr.NANot Available1969DEVELOPEDCultivarFrom the United States. Selections made by J.L. Creech and W.L. Ackerman at the United States Plant Introduction Station, Glenn Dale, MD. Numbered June 25, 1968. Selection from an F1 progeny resulting from a cross of Ilex cornuta, PI65860 x I. ciliospinosa, PI78144 made in 1952. 'Howard Dorsett'. Male evergreen shrub, compact, pyramidal. Leaves medium green, glossy, spinose, oblong lanceolate, 3 to 5 cm long, 1 1/2 to 2 cm wide, mostly 9 spines. New growth pinkish, turning green. Considered good companion plant to 'William Cowgill'.2107846PI 331204 NA 31492
32PI 331206 NA 31491'Harry Gunning'Ilex hybr.NANot Available1969DEVELOPEDCultivarFrom the United States. Selections made by J.L. Creech and W.L. Ackerman at the United States Plant Introduction Station, Glenn Dale, MD. Numbered June 25, 1968. Selection from an F1 progeny resulting from a cross of Ilex cornuta, PI65860 x I. ciliospinosa, PI78144 made in 1952. 'Harry Gunning'. Male evergreen shrub, compact, globular. Leaves medium green, oblong lanceolate, 3 to 4 1/2 cm long, 1 1/4 to 1 3/4 cm wide, mostly 7 spines.2107847PI 331206 NA 31491
33PI 560965 NA 31330'Coronet'Ilex hybr.District of Columbia, United StatesNANot Available1969DEVELOPEDCultivarDonated by Eisenbeiss, G., U.S. National Arboretum, USDA-ARS, 3501 New York Avenue, NE, Washington, D.C. 20002, United States. Received April 2, 1992. Originated from cross made 1963 at U.S. National Arboretum by W.F. Kosar (pedigree K62-19-3). 'Coronet'. Ilex [(aquifolium/cornuta) 'Nellie R. Stevens'/I. ciliospinosa/(I. /aquipernyi)]. Large evergreen shrub to 15'. Leaves soft, spiny. Female. Fruit red. Hardiness zone 7 on USDA Plant Hardiness Zone Map (1990). International Ilex cultivar registration 12-91. Selected by W.F. Kosar and named by G.K. Eisenbeiss. Cultivar. Plant. Had been tentatively named 'Self-fruitful Lady' by Kosar as of January 1971, per the sheet titled "Holly and Magnolia Hybrids Tentatively Named by W. F. Kosar as of Jan 1971"1455937PI 560965 NA 31330
34PI 378149 NA 29738'Sparkleberry'Ilex hybr.District of Columbia, United StatesNANot Available1967DEVELOPED1961CultivarFull name as recorded at the U.S. National Arboretum: Ilex 'Sparkleberry'. From United States. Plant selections made in 1961 by W.F. Kosar at United States National Arboretum, Washington, D.C. Received January 30, 1973. 'Sparkleberry'. A female selection from progeny of the cross I. serrata x I. verticillata. Decdiuous, upright shrub. Leaves intermediate in character between the parent species. Fruit superior in size, gloss, and persistance to I. serrata and more productive than I. verticillata. Hardiness 5A on USDA Hardiness Zone Map.2107850PI 378149 NA 29738
35PI 422217 NA 29739'Apollo'Ilex hybr.District of Columbia, United StatesNANot Available1967DEVELOPED1961CultivarFull name as recorded at the U.S. National Arboretum: Ilex 'Apollo'. From United States. Plant selections made by W.F. Kosar at the United States National Arboretum, Washington D.C. Numbered February 17, 1978. Deciduous, upright shrub. Leaves intermediate between parent species. Male selection as a pollinator for PI378149 to which it is a sibling. Originated in 1961 from NA 11876 (I. serrata) x GE 685 (I. verticillata). Hardiness 5A on USDA Plant Hardiness Zone map.2107853PI 422217 NA 29739
36PI 319168Ilex aquifolium L. Krym, Ukraine Historic1967COLLECTED1711101PI 319168
37PI 319169Ilex colchica Pojark. Krym, Ukraine Historic1967COLLECTED1711102PI 319169
38PI 319170Ilex cornuta Lindl. & Paxton Krym, Ukraine Historic1967COLLECTED1711103PI 319170
39PI 319171Ilex latifolia Thunb. Krym, Ukraine Historic1967COLLECTED1711104PI 319171
40PI 318660Ilex insignis Hook. f. India Historic1967COLLECTED1709553PI 318660
41PI 318661Ilex wightiana Wall. ex Wight India Historic1967COLLECTED1709554PI 318661
42PI 318652Ilex aquifolium L. Hordaland, Norway Historic1967COLLECTED1709547PI 318652
43PI 318549Col. No. K-13Ilex crenata Thunb. Jeju-teukbyeoljachido, Korea, South Historic1967COLLECTEDSouth slope, Mt. Halla, Cheju Island, moist open area1200Wild materialSmall leaved, evergreen, dwarf shrub with small white flowers. Male plant. Plants. 10 1237106PI 318549
44PI 317234Col. No. K-442Ilex crenata Thunb. Jeju-teukbyeoljachido, Korea, South Historic1966COLLECTEDOpen alpine meadow, Mt. Halla, Cheju Island14001708746PI 317234
45PI 317235Col. No. K-426Ilex crenata Thunb. Jeju-teukbyeoljachido, Korea, South Historic1966COLLECTEDAlong trail up Mt. Halla, Cheju Island7001708747PI 317235
46PI 317035No. 733Ilex canariensis Poir. Canarias, Spain Historic1966COLLECTED1708358PI 317035
47PI 317040No. 327Ilex mitis (L.) Radlk. South Africa Historic1966COLLECTED1708363PI 317040
48PI 316703Col. No. K-249Ilex macropoda Miq. Korea, South Historic1966COLLECTEDIn open, along trail on Mt. Sok Ni4501708213PI 316703
49PI 316704Col. No. K-274Ilex macropoda Miq. Korea, South Historic1966COLLECTEDAlong road to Mt. Tok-Yu, Ko Chun1708214PI 316704
50PI 316603Col. No. K-133Ilex macropoda Miq. Korea, South Historic1966COLLECTEDAlong trail up Mt. Chiri, Cholla Namdo11001708088PI 316603
51PI 316588'High Light'Ilex crenata Thunb. District of Columbia, United States Historic1966COLLECTED1708072PI 316588
52PI 315894Ilex pedunculosa Miq. Japan Historic1966COLLECTEDKaseda, Kyusiu Island1707714PI 315894
53PI 315213Ilex colchica Pojark. Russian Federation Historic1966COLLECTED1707457PI 315213
54PI 329155 NA 28323'Tanager'Ilex hybr.NANot Available1966CultivarFrom the United States. Selections made at the National Arboretum, United States Department of Agriculture, Washington, D.C. Received Apr. 15, 1968. F2 selections from I. myrtifolia x I. opaca. Plants slow growing, compact, fine textured; leaves spinose, 1/2 by 1 3/4 inches; fruit red. 'Tanager'.2107844PI 329155 NA 28323
55PI 313116'Mariesii'Ilex crenata Thunb. United Kingdom Historic1966COLLECTED1706182PI 313116
56PI 311717Ilex sikkimensis Kurz United Kingdom Historic1966COLLECTED1705494PI 311717
57PI 311301Ilex intricata Hook. f. India Historic1966COLLECTED1705293PI 311301
58PI 578192'Geisha'Ilex crenata Thunb. District of Columbia, United States Historic1966DONATED03/23/1994CultivarDonated by G. Eisenbeiss, USDA-ARS, National Arboretum, 3501 New York Ave., N.D., Washington, D.C. 20002, United States. Received [PI numbered] 3/23/1994. 'Geisha'. Female parent PI 231948, NA 10815 a yellow fruited plant from Japan. Male parent NA 10822 a F2 backcross of a male seedling of NA 10815, back crossed to 10815. Evergreen shrub with extremely small convex leaves and yellow fruit.1473163PI 578192
59PI 308634Ilex asprella (Hook. & Arn.) Champ. ex Benth. Hong Kong Historic1965COLLECTED1704344PI 308634
60PI 308635Ilex pubescens Hook. & Arn. Hong Kong Historic1965COLLECTED1704345PI 308635
61PI 308636Ilex viridis Champ. ex Benth. Hong Kong Historic1965COLLECTED1704346PI 308636
62PI 307274Col. No. 122Ilex hookeri King West Bengal, India Historic1965COLLECTEDBetween Phalut and Samanden28601702786PI 307274
63PI 307275Col. No. 208Ilex insignis Hook. f. West Bengal, India Historic1965COLLECTED1702787PI 307275
64PI 307276Col. No. 176Ilex intricata Hook. f. West Bengal, India Historic1965COLLECTEDMt. Tonglu, along trail from Batasi24081702788PI 307276
65PI 307277Col. No. 209Ilex wightiana Wall. ex Wight West Bengal, India Historic1965COLLECTED1702789PI 307277
66PI 307278Col. No. 282Ilex wightiana Wall. ex Wight West Bengal, India Historic1965COLLECTEDDarjeeling1702790PI 307278
67PI 306368366, 499Ilex aquifolium L. Germany Historic1965COLLECTED1702235PI 306368
68PI 305310Ilex aquifolium L. Denmark Historic1965COLLECTED1701265PI 305310
69PI 305311Ilex aquifolium L. Denmark Historic1965COLLECTED1701266PI 305311
70PI 305312Ilex aquifolium L. Denmark Historic1965COLLECTED1701268PI 305312
71PI 305313'Pyramidalis'Ilex aquifolium L. Denmark Historic1965COLLECTED1701267PI 305313
72PI 304944Ilex crenata Thunb. Hirosima, Japan Historic1965COLLECTED1701086PI 304944
73PI 304945Ilex crenata Thunb. Hirosima, Japan Historic1965COLLECTED1701087PI 304945
74PI 304946Ilex purpurea Hassk. Kagosima, Japan Historic1965COLLECTED1701088PI 304946
75PI 304947Ilex rotunda Thunb. Hirosima, Japan Historic1965COLLECTED1701089PI 304947
76PI 260386 NA 25704NA 25704Ilex crenata Thunb. Honshu, JapanNANot Available1964COLLECTEDPRE 09/09/1959Aomori Prefecture: Mutsu-shi.Wild materialReceived as a single seedling, inocrrectly identified as I. crenata var. paludosa. From USDA Plant Inventory No. 167: "260384 to 260393. From Japan. Seeds presented by the Government Forest Experiment Station, Okidate, Aomori. Received September 9, 1959. [...] 260385 and 260386. Ilex crenata var. paludosa Thunb. Aquifoliaceae. [...] 260386. Province Mutsu." [This accession does not display the low and spreading (or low and creeping) habit of I. crenata var. paludosa found in other individuals. Instead, the habit is vertical, significantly taller than wide, yet open (not narrowly fastigiate). Both remaining plants of this accession have this appearance (which makes sense, since they belong to a single seedling clone). The closely-ranked foliage is also interesting, being more prominently and sharply toothed than typical Ilex crenata. Male.2191651PI 260386 NA 25704
77PI 299436Ilex asprella (Hook. & Arn.) Champ. ex Benth. Hong Kong Historic1964COLLECTED1677912PI 299436
78PI 299437Ilex cinerea Champ. ex Benth. Hong Kong Historic1964COLLECTED1677913PI 299437
79PI 299438Ilex graciliflora Champ. ex Benth. Hong Kong Historic1964COLLECTED1677914PI 299438
80PI 299439Ilex memecylifolia Champ. ex Benth. Hong Kong Historic1964COLLECTED1677915PI 299439
81PI 299440Ilex pubescens Hook. & Arn. Hong Kong Historic1964COLLECTED1677916PI 299440
82PI 299441Ilex rotunda Thunb. Hong Kong Historic1964COLLECTED1677917PI 299441
83PI 299399Ilex mitis (L.) Radlk. South Africa Historic1964COLLECTED1677895PI 299399
84PI 298031Ilex mitis (L.) Radlk. South Africa Historic1964COLLECTED1682694PI 298031
85PI 296985No. 1Ilex centrochinensis S. Y. Hu Japan Historic1964COLLECTED1681832PI 296985
86PI 296986No. 2Ilex centrochinensis S. Y. Hu Japan Historic1964COLLECTED1681833PI 296986
87PI 296020Ilex macropoda Miq. Iwate, Japan Historic1964COLLECTED1681335PI 296020
88PI 292361No. 372Ilex mitis (L.) Radlk. South Africa Historic1963COLLECTED1695403PI 292361
89PI 291938Ilex verticillata (L.) A. Gray Ontario, Canada Historic1963COLLECTEDHamilton1694777PI 291938
90PI 291939Ilex verticillata (L.) A. Gray Canada Historic1963COLLECTEDNorthumberland County, Colborne1694778PI 291939
91PI 291941No. 12362Ilex mucronata (L.) M. Powell et al. Ontario, Canada Historic1963COLLECTEDBorthwick Road, 6.4km east of Ramsayville1694780PI 291941
92PI 289935Ilex sp. Nepal Historic1963COLLECTED1692350PI 289935
93PI 289743'Golden Gem'Ilex crenata Thunb. Netherlands Historic1963COLLECTED1692248PI 289743
94PI 288703Ilex aquifolium L. Japan Historic1963COLLECTED1691441PI 288703
95PI 288704Ilex centrochinensis S. Y. Hu Japan Historic1963COLLECTED1691442PI 288704
96PI 286159Ilex macropoda Miq. Japan Historic1963COLLECTED1689099PI 286159
97PI 560963 NA 28358'Venus'Ilex hybr.District of Columbia, United StatesNANot Available1963DEVELOPEDCultivarDonated by Eisenbeiss, G., U.S. National Arboretum, USDA-ARS, 3501 New York Avenue, NE, Washington, D.C. 20002, United States. Received April 2, 1992. Originated from cross made 1963 at U.S. National Arboretum by W.F. Kosar (pedigree K63-4-2). 'Venus'. Ilex [(aquifolium/cornuta) 'Nellie R. Stevens'/I. latifolia]. Large evergreen shrub to 18'. Numerous small spines on leaf margin. Female. Fruit red. Hardiness zone 7 on USDA Plant Hardiness Zone Map (1990). International Ilex cultivar registration 8-91. Selected and named by F.S. Santamour, Jr. and G.K. Eisenbeiss. Cultivar. Plant.1455935PI 560963 NA 28358
98PI 560964 NA 28313'Adonis'Ilex hybr.District of Columbia, United StatesNANot Available1963DEVELOPEDCultivarDonated by Eisenbeiss, G., U.S. National Arboretum, USDA-ARS, 3501 New York Avenue, NE, Washington, D.C. 20002, United States. Received April 2, 1992. Originated from cross made 1963 at U.S. National Arboretum by W.F. Kosar (pedigree K63-4-1). 'Adonis'. Ilex [(aquifolium/cornuta) 'Nellie R. Stevens'/I. latifolia]. Large evergreen shrub to 18'. Leaves large with numerous small spines on leaf margin. Male. Hardiness zone 7 on USDA Plant Hardiness Zone Map (1990). International Ilex cultivar registration 9-91. Selected and named by F.S. Santamour, Jr. and G.K. Eisenbeiss. Cultivar. Plant.1455936PI 560964 NA 28313
99PI 285365Col. No. 1272Ilex excelsa (Wall.) Hook. f. Nepal Historic1962COLLECTEDAlong streambed above Godavari18061688818PI 285365
100PI 285445Col. No. 1154Ilex dipyrena Wall. Nepal Historic1962COLLECTEDIn forest with rhododendrons, Mt. Furkai25591688895PI 285445
101PI 285469Col. No. 1234Ilex sp. Nepal Historic1962COLLECTEDRhododendron forest above Mardi Khola24081688918PI 285469
102PI 284783Ilex paraguariensis A. St.-Hil. Australia Historic1962COLLECTED1687948PI 284783
103PI 282697Ilex asprella (Hook. & Arn.) Champ. ex Benth. Hong Kong Historic1962COLLECTED1686697PI 282697
104PI 282698Ilex hanceana Maxim. Hong Kong Historic1962COLLECTED1686698PI 282698
105PI 282478Ilex mitis (L.) Radlk. South Africa Historic1962COLLECTED1686569PI 282478
106PI 282427 NA 19791'Agena'Ilex ×koehneana Loes. District of Columbia, United StatesNANot Available1962DEVELOPEDCultivarFrom New Zealand. Plants purchased from Duncan and Davies, Westown, New Plymouth. Received July 30, 1962. [Later reidentified as Ilex x koehneana, selected and introduced by the U.S. National Arboretum as 'Agena'.]1209826PI 282427 NA 19791
107PI 279951Ilex centrochinensis S. Y. Hu Japan Historic1962COLLECTED1685028PI 279951
108PI 279952DEIP No. 21.274Ilex paraguariensis A. St.-Hil. Argentina Historic1962COLLECTED1685029PI 279952
109PI 280057'Madame Briot'Ilex aquifolium L. United Kingdom Historic1962COLLECTED1685065PI 280057
110PI 280058'Longifolia'Ilex aquifolium L. United Kingdom Historic1962COLLECTED1685066PI 280058
111PI 279006JC0461Ilex crenata Thunb. Japan Historic1962COLLECTED1700507PI 279006
112PI 276341JC0461Ilex rotunda Thunb. Maryland, United States Historic1961DONATED08/17/19611206711PI 276341
113PI 272055 NA 17022PI 272055Ilex serrata Thunb. NANot Available1961Cultivated materialFrom the record of NA 17022 at the U.S. National Arboretum: "From Japan. Seeds purchased from P. H. Kubota, Nikko Tochigi Prefecture. Received March 3, 1961. Collected from cultivated plants in Angyo, Saitama Prefecture. Comments: 9 seedlings flowered 3 males, 6 females; 1 female yellow-fruited; the remaining red-fruited. Only 3 listed in locations remain NA 17022-1 named 'Sundrops' 1991. NA 17022-1 reaccessioned to NA 62386 (3/91) to be named and released by G. Eisenbeiss. See NA 62386 'Sundrops' & HSA Reg. 1991. Name changed to Ilex serrata - not all seedlings bear white/yellow fruit so name is inappropriate, and according to Eisenbeiss's notes, only one NA 17022 seedling actually bore yellow fruit and became 'Sundrops', so presumably the remaining fraction are red-fruited or male. They should, however, at least carry the recessive gene for yellow/pale fruit. -SBL"2158576PI 272055 NA 17022
114PI 560967 NA 62386'Sundrops'Ilex serrata Thunb. District of Columbia, United StatesNANot Available1961DEVELOPEDCultivarDonated by Eisenbeiss, G., U.S. National Arboretum, USDA-ARS, 3501 New York Avenue, NE, Washington, D.C. 20002, United States. Received April 2, 1992. Seed from Japan 1961 grown at U.S. National Arboretum. Seed was originally purchased from P.H. Kubota, Nikko Tochigi Prefecture, Japan was was collected from cultivated plants in Angyo Saitama Prefecture. 'Sundrops'. Selection of PI272055, NA 17022. Deciduous shrub to 8' tall and 15' wide. Female. Fruit yellow. Hardiness zone 5 on USDA Plant Hardiness Zone Map (1990). International Ilex cultivar registration 14-91. Cultivar. Plant. [Information from the accession number of the parent lot, NA 17022: From Japan. Seeds purchased [as Ilex serrata forma leucocarpa] from P. H. Kubota, Nikko Tochigi Prefecture. Received March 3, 1961. Collected from cultivated plants in Angyo, Saitama Prefecture. Comments: 9 seedlings flowered 3 males, 6 females; 1 female yellow-fruited; the remaining red-fruited. Only 3 listed in locations remain NA 17022-1; named 'Sundrops' 1991. NA 17022-1 reaccessioned to NA 62386 (3/91) to be named and released by G. Eisenbeiss. See NA 62386 'Sundrops' & HSA Reg. 1991.]1455939PI 560967 NA 62386
115PI 560966 NA 28362'Miniature'Ilex hybr.District of Columbia, United StatesNANot Available1961DEVELOPEDCultivarDonated by Eisenbeiss, G., U.S. National Arboretum, USDA-ARS, 3501 New York Avenue, NE, Washington, D.C. 20002, United States. Received April 2, 1992. Originated from cross made 1961 at U.S. National Arboretum by W.F. Kosar Interspecific Hybrid (pedigree K61-15-4). 'Miniature'. 7075 = Ilex [(aquifolium/cornuta) 'Nellie R. Stevens'/NA 9661 = I. pernyi]. Evergreen shrub to 8'. Leaves small, strongly spined. Female. Fruit red. Hardiness zone 7 on USDA Plant Hardiness Zone Map (1990). International Ilex cultivar registration 13-91. Selected and named by W.F. Kosar. Cultivar. Plant.1455938PI 560966 NA 28362
116PI 269620 NA 18381'Ajax'Ilex ×koehneana Loes. District of Columbia, United StatesNANot Available1960DEVELOPEDCultivarOriginal plant grown from a seedling (30 cm in height) of unknown and disputed botanical status sent to the Arboretum for identification in 1960. Selected and named by S.F. Santamour, Jr. and G.K. Eisenbeiss. Fast-growing pyramidal evergreen tree to 35 feet in height. Leaves broadly lanceolate to 5 inches long, margins continuous with small uniform spines. Selected as male pollinator of similar size and habit for Agena. Hardiness is Zone 7. PI 269620:From Great Britain. Plants purchased from H.G. Hillier & Sons, Winchester, Hants, England. Received Nov. 14, 1960.1202833PI 269620 NA 18381
117PI 329153 NA 28261'Elegance'Ilex hybr.NANot Available1960CultivarFrom the United States. Selections made at the National Arboretum, United States Department of Agriculture, Washington, D.C. Received Apr. 15, 1968. F1 selections from I. integra x I. pernyi. Plants single-stemmed, conical, compact; leaves small, evergreen. 'Elegance'. Fruit red, colors early.2107842PI 329153 NA 28261
118PI 329154 NA 28260'Accent'Ilex hybr.NANot Available1960CultivarFrom the United States. Selections made at the National Arboretum, United States Department of Agriculture, Washington, D.C. Received Apr. 15, 1968. F1 selections from I. integra x I. pernyi. Plants single-stemmed, conical, compact; leaves small, evergreen. 'Accent'. Excellent accent plant in landscaping. Good pollinator for 'Elegance'.2107843PI 329154 NA 28260
119PI 371685 NA 28341'Ruby'Ilex ×koehneana Loes. NANot Available1960DEVELOPEDCultivarPI 371685: From United States. Plants developed by the National Arboretum, Plant Science Research Division, Washington, D.C. Received March 13, 1972. 'Ruby'. Female selection from F1 population of hybrid I. aquifolium x I. latifolia, made by William F. Kosar. Vigorous, conical tree to 6.5 m tall. More hardy and more tolerant to adverse exposure and soil than parent species. Leaves evergreen, 15 to 18 cm long, intermediate in number and length of spines between parent species. Fruits bright red, borne on short pedicels, intermediate in size between those of parents.1278576PI 371685 NA 28341
120PI 371686 NA 28342'Jade'Ilex ×koehneana Loes. NANot Available1960DEVELOPEDCultivarFrom United States. Plants developed by the National Arboretum Plant Science Research Division, Washington, D.C. Received March 13, 1972. 'Jade'. Male selection from F1 population hybrid between I. aquifolium x I. latifolia and a brother seedling of PI371685. Very similar to 'Ruby' in foliage, habit of growth, and hardiness and is being introduced as a pollinator for 'Ruby'.1278577PI 371686 NA 28342
121PI 267253'Media Picta'Ilex aquifolium L. Scotland, United Kingdom Historic1960DEVELOPEDCultivar1201272PI 267253
122PI 267255'Recurva'Ilex aquifolium L. Scotland, United Kingdom Historic1960DEVELOPEDCultivar1201273PI 267255
123PI 267257'Whittingtonensis'Ilex aquifolium L. Scotland, United Kingdom Historic1960DEVELOPEDCultivar1201274PI 267257
124PI 267259'Ferox Aurea'Ilex aquifolium L. Scotland, United Kingdom Historic1960DONATED06/14/1960Cultivar1201275PI 267259
125PI 267261'Ferox'Ilex aquifolium L. Scotland, United Kingdom Historic1960DEVELOPEDCultivar1201276PI 267261
126PI 267263 NA 15342'Pendula'Ilex aquifolium L. Scotland, United KingdomNANot Available1960DEVELOPEDCultivar1201277PI 267263 NA 15342
127PI 267266'Crassifolia'Ilex aquifolium L. Scotland, United Kingdom Historic1960DONATED06/14/1960Cultivar1201278PI 267266
128PI 267267 NA 15351NA 15351Ilex crenata Thunb. Scotland, United KingdomNANot Available1960DONATED06/14/1960CloneReceived as Ilex crenata f. latifolia major. From the record of NA15351 at the U.S. National Arboretum: 9/86: T. R. Dudley changed name from f. latifolia major to cv. Major; 3/94: TR Dudley & G. Eisenbeiss noted that the cv. name Major has been applied to several clones. We are unsure of what we have, so we reduced the name to the species. (see Ilex crenata checklist).1201279PI 267267 NA 15351
129PI 263648Ilex crenata Thunb. Hukuoka, Japan Historic1960DONATED02/18/19601568064PI 263648
130PI 263649'Convexa'Ilex crenata Thunb. Hukuoka, Japan Historic1960DONATED02/18/19601568065PI 263649
131PI 263651Ilex integra Thunb. Hukuoka, Japan Historic1960DONATED02/18/19601568066PI 263651
132PI 263652Ilex latifolia Thunb. Hukuoka, Japan Historic1960DONATED02/18/19601568067PI 263652
133PI 263653Ilex pedunculosa Miq. Hukuoka, Japan Historic1960DONATED02/18/19601568068PI 263653
134PI 263654Ilex rotunda Thunb. Hukuoka, Japan Historic1960DONATED02/18/19601568069PI 263654
135PI 263655Ilex serrata Thunb. Hukuoka, Japan Historic1960DONATED02/18/19601568070PI 263655
136PI 262783Ilex latifolia Thunb. Saitama, Japan Historic1960DONATED01/06/19601567351PI 262783
137PI 578191 NA 28273'Scepter'Ilex hybr.District of Columbia, United StatesNANot Available1960DONATED03/23/1994CultivarControlled cross [using mixed pollen] and selection made by W. F. Kosar at the U.S. National Arboretum. Named and introduced by G. Eisenbeiss, USDA-ARS, National Arboretum, 3501 New York Ave., N.E., Washington, D.C. 20002, United States. NPGS received 3/23/1994. 'Scepter'. Ilex [integra x (x altaclerensis 'Hodginsii')]. Red fruited evergreen tree to 30 feet with soft, spineless to weakly spined leaves. [Kosar had given this clone the internal, manuscript name "Amazon".]1473162PI 578191 NA 28273
138PI 261745'Brevipedunclata'Ilex sugerokii Maxim. Tôkyô, Japan Historic1959DONATED12/09/19591566725PI 261745
139PI 261441Ilex perado Aiton Maryland, United States Historic1959DONATED11/25/19591565884PI 261441
140PI 261224'Bicolor'Ilex aquifolium L. England, United Kingdom Historic1959DONATED11/12/19591565713PI 261224
141PI 261225'Latispina Minor'Ilex aquifolium L. England, United Kingdom Historic1959DONATED11/12/19591565714PI 261225
142PI 261226'N.E. Barnes'Ilex aquifolium L. England, United Kingdom Historic1959DONATED11/12/19591565715PI 261226
143PI 261227'Recurva'Ilex aquifolium L. England, United Kingdom Historic1959DONATED11/12/19591565716PI 261227
144PI 261228'Variegata'Ilex aquifolium L. England, United Kingdom Historic1959DONATED11/12/19591565717PI 261228
145PI 261229Ilex cassine L. England, United Kingdom Historic1959DONATED11/12/19591565718PI 261229
146PI 261230Ilex corallina Franch. England, United Kingdom Historic1959DONATED11/12/19591565719PI 261230
147PI 261231Ilex dipyrena Wall. England, United Kingdom Historic1959DONATED11/12/19591565720PI 261231
148PI 261233'Sclerophylla'Ilex fargesii Franch. England, United Kingdom Historic1959DONATED11/12/19591565721PI 261233
149PI 261234Ilex hookeri King England, United Kingdom Historic1959DONATED11/12/19591565722PI 261234
150PI 261235Ilex fargesii Franch. subsp. melanotricha (Merr.) S. Andrews England, United Kingdom Historic1959DONATED11/12/19591565723PI 261235
151PI 261236Ilex yunnanensis Franch. England, United Kingdom Historic1959DONATED11/12/19591565724PI 261236
152PI 261237'Forest 24826'Ilex sp.England, United Kingdom Historic1959DONATED11/12/19591565725PI 261237
153PI 261216Ilex fargesii Franch. subsp. melanotricha (Merr.) S. Andrews England, United Kingdom Historic1959DONATED11/10/19591565709PI 261216
154PI 260749Ilex cornuta Lindl. & Paxton Maryland, United States Historic1959DONATED10/13/19591563455PI 260749
155PI 260385Ilex crenata Thunb. var. paludosa (Nakai) H. Hara Honshu, Japan Historic1959DONATED09/09/19591563168PI 260385
156PI 260386Ilex crenata Thunb. Honshu, JapanNANot Available1959COLLECTEDPRE 09/09/1959Aomori Prefecture: Mutsu-shi.Wild materialFrom the record of NA 25704 at the U.S. National Arboretum: From USDA Plant Inventory No. 167: "260384 to 260393. From Japan. Seeds presented by the Government Forest Experiment Station, Okidate, Aomori. Received September 9, 1959. [...] 260385 and 260386. Ilex crenata var. paludosa Thunb. Aquifoliaceae. [...] 260386. Province Mutsu." [This accession does not display the low and spreading (or low and creeping) habit of I. crenata var. paludosa found in other individuals. Instead, the habit is vertical, significantly taller than wide, yet open (not narrowly fastigiate). Both remaining plants of this accession have this appearance. -SBL2158600PI 260386
157PI 260387Ilex macropoda Miq. Honshu, Japan Historic1959DONATED09/09/19591563169PI 260387
158PI 260388Ilex leucoclada (Maxim.) Makino Honshu, Japan Historic1959DONATED09/09/19591563170PI 260388
159PI 260389Ilex sugerokii Maxim. Honshu, Japan Historic1959DONATED09/09/19591563171PI 260389
160PI 235574 NA 12679PI 235574Ilex triflora Blume var. kanehirae (Yamam.) S. Y. Hu NANot Available1959Cultivated materialReceived as Ilex mutchagara at the U.S. National Arboretum. From Japan. Plants collected by J.L. Creech, Agricultural Explorer, Plant Introduction Section, Horticultural Crops Research Branch, Beltsville, MD. Received Nov. 14, 1956. Collected in Nov., 1956. Should be hardy in Maryland. PI book: "Col. No. 578. Kyoto University Botanic Garden, Kyoto. Evergreen tree to 20 feet high: leaves ovate-lanceolate; fruits black."2158542PI 235574 NA 12679
161PI 255344Ilex aquifolium L. Slovenia Historic1959DONATED02/06/19591558166PI 255344
162PI 254592Ilex cassine L. England, United Kingdom Historic1958DONATED12/12/19581195485PI 254592
163PI 422216'September Gem'Ilex hybr.NANot Available19582107852PI 422216
164PI 231705 NA 10821'Watereriana'Ilex aquifolium L. England, United KingdomNANot Available1957DONATED03/14/1956CultivarReceived as grafted plants.1184010PI 231705 NA 10821
165PI 237879Ilex rotunda Thunb. Japan Historic1957COLLECTEDWild material1186374PI 237879
166PI 236233 NA 25701NakadaIlex crenata Thunb. JapanNANot Available1957COLLECTEDCultivarFrom Japan. Plants collected by J.L. Creech, agricultural explorer, Plant Introduction Section, Horticultural Crops Research Branch, Beltsville, MD. Received Jan. 3, 1957. Horticultural varieties of plants purchased from Nakada Nursery, Angyo, Kawaguchi, Saitamaken.1185655PI 236233 NA 25701
167PI 231948Ilex crenata Thunb. District of Columbia, United States Historic1956DEVELOPED1968Cultivar1492198PI 231948
168PI 267824 NA 12202'Lydia Morris'Ilex hybr.NANot Available1952CultivarFrom the United States. Plants growing at the National Arboretum, Washington, D.C. Numbered August 15, 1960. 'Lydia Morris' (I. cornuta 'Burfordi' x I. pernyi). Pistillate hybrid; plants narrowly pyramidal to about six feet high; leaves lustrous, intermediate; berries large, red on short pedicels. Ilex hybrid F1 crosses made by Dr. Henry T. Skinner, Spring 1948 at Morris Arboretum. [Note: the parent accession was numbered later as NA 21219. -SBL]2107833PI 267824 NA 12202
169PI 267825 NA 12201'John T. Morris'Ilex hybr.District of Columbia, United StatesNANot Available1952DONATED08/15/1960CultivarFrom the United States. Plants growing at the National Arboretum, Washington, D.C. Numbered August 15, 1960. 'John T. Morris'. (I. cornuta 'Burfordi' x I. pernyi). Staminate hybrid with leaves and growth habit similar to 'Lydia Morris', a sister seedling. Ilex hybrid F1 crosses made by Dr. Henry T. Skinner, Spring 1948 at Morris Arboretum. [Note: the parent accession was numbered later as NA 21219. -SBL]1201580PI 267825 NA 12201
170PI 198275B-43193Ilex aquifolium L. Maryland, United States Historic1951COLLECTED1675898PI 198275
171PI 198276B-43194Ilex aquifolium L. Maryland, United States Historic1951COLLECTED1675899PI 198276
172PI 198277B-43195Ilex cornuta Lindl. & Paxton Maryland, United States Historic1951COLLECTED1675900PI 198277
173PI 198278B-10909Ilex cornuta Lindl. & Paxton Maryland, United States Historic1951COLLECTED1675901PI 198278
174PI 198279B-10909AIlex cornuta Lindl. & Paxton Maryland, United States Historic1951COLLECTED1675902PI 198279
175PI 198280Ilex sp.Maryland, United States Historic1951COLLECTED1675903PI 198280
176PI 195133Ilex cornuta Lindl. & Paxton Maryland, United States Historic1951COLLECTED1671868PI 195133
177PI 148016Ilex sp.Virginia, United States Historic1944COLLECTEDBennet's Creek1881606PI 148016
178PI 148017Ilex sp.Virginia, United States Historic1944COLLECTEDBennet's Creek1881607PI 148017
179PI 146599Ilex sp.Amazonas, Brazil Historic1943COLLECTED05/08/1943Porto Velho1879927PI 146599
180PI 143795 NA 1848'Rotunda'Ilex cornuta Lindl. & Paxton Louisiana, United StatesNANot Available1942COLLECTEDCultivarFrom Louisiana. Cuttings presented by E.A. McIlhenny, Avery Island. Received February 6, 1942. PI143795, A variety that develops a compact, rounded habit. References: x 10/10/41 E. A. McIlhenny to RA Young. x 11/1/41 RAY to EAM. x 12/31/41 RAY to EAM.1869732PI 143795 NA 1848
181PI 141022No. 6347Ilex sp.Māzandarān, Iran Historic1941COLLECTED07/23/1940Dimalu1865646PI 141022
182PI 139217No. 5293Ilex coriacea (Pursh) Chapm. Georgia, United States Historic1940COLLECTED08/24/1940Homerville1862378PI 139217
183PI 97994 NA 280'Camelliifolia'Ilex ×altaclerensis (Loudon) Dallim. NetherlandsNANot Available1939COLLECTEDCultivarFrom the Netherlands. Plants purchased from B. Ruys, Ltd., Royal Moerheim Nurseries, Dedemsvaart. Received March 16, 1932. PI97994, Leaves elliptic or oblong, acuminate, dark olive green, very glossy, entire or with a few spines near the apex.1784624PI 97994 NA 280
184PI 107438 NA 25(seed of 'Aurifodina')Ilex aquifolium L. New York, United StatesNANot Available1938COLLECTEDCultivated materialFrom New York. Seeds presented by J.B. Murray, Brooklyn. Received December 20, 1934. PI107438; Variety Aurifodina.1796806PI 107438 NA 25
185PI 107443 NA 29(seed of 'Fructu Luteo')Ilex aquifolium L. New York, United StatesNANot Available1938COLLECTEDCultivated materialFrom New York. Seeds presented by J.B. Murray, Brooklyn. Received December 20, 1934. PI107443; Variety Fructu-luteo.1796811PI 107443 NA 29
186PI 207438NA 25Ilex aquifolium L. Colombia Historic1938DONATED10/19/1938Cultivated material1173508PI 207438
187PI 207439NA 26Ilex aquifolium L. Colombia Historic1938DONATED10/19/1938Cultivated material1173509PI 207439
188PI 207440NA 27Ilex aquifolium L. Colombia Historic1938DONATED10/19/1938Cultivated material1173510PI 207440
189PI 130082No. 7826Ilex sp.Yunnan Sheng, China Historic1938COLLECTEDDokerla, Atuntze30001846009PI 130082
190PI 130083No. 7831Ilex sp.Yunnan Sheng, China Historic1938COLLECTEDDokerla, Atuntze31001846010PI 130083
191PI 130084No. 10610Ilex sp.Yunnan Sheng, China Historic1938COLLECTEDMount Kaaperpu, Atuntze30001846011PI 130084
192PI 130085No. 13536Ilex sp.Yunnan Sheng, China Historic1938COLLECTEDHaba, Chungtien34001846012PI 130085
193PI 130086No. 14283Ilex sp.Yunnan Sheng, China Historic1938COLLECTEDKulu, Muli32001846013PI 130086
194PI 130087No. 14921Ilex sp.Yunnan Sheng, China Historic1938COLLECTEDPaitih, Chungtien27001846014PI 130087
195PI 129234No. 1278Ilex sp.Sichuan Sheng, China Historic1938COLLECTED10/1937Mount Omei6001845272PI 129234
196PI 129235No. 1285Ilex sp.Sichuan Sheng, China Historic1938COLLECTED10/1937Mount Omei60001845273PI 129235
197PI 129236No. 1288Ilex sp.Sichuan Sheng, China Historic1938COLLECTED10/1937Mount Omei between 900m and 1000m altitude1845274PI 129236
198PI 129237No. 1313Ilex sp.Sichuan Sheng, China Historic1938COLLECTED11/1937Tien Chuan Hsien23001845275PI 129237
199PI 129238No. 1349Ilex sp.Sichuan Sheng, China Historic1938COLLECTED11/1937O-pie Hsien17001845276PI 129238
200PI 129239No. 1373Ilex sp.Sichuan Sheng, China Historic1938COLLECTED10/1937Mount Omei18001845277PI 129239
201PI 129240No. 1396Ilex sp.Sichuan Sheng, China Historic1938COLLECTED11/1937Tien Chuan Hsien5001845278PI 129240
202PI 124708Ilex pedunculosa Miq. China Historic1937COLLECTED1839275PI 124708
203PI 124250Ilex hookeri King India Historic1937COLLECTED1838086PI 124250
204PI 124251Ilex insignis Hook. f. India Historic1937COLLECTED1838087PI 124251
205PI 122095No. 1919Ilex insignis Hook. f. West Bengal, India Historic1937COLLECTED12/1936Darjeeling1833020PI 122095
206PI 122096No. 1920Ilex sp.West Bengal, India Historic1937COLLECTED12/1936Darjeeling1833021PI 122096
207PI 122673No. 2401Ilex sp.Trabzon, Turkey Historic1937COLLECTED08/12/193623km east of Trabzon1834157PI 122673
208PI 117060Ilex geniculata Maxim. Japan Historic1936COLLECTED1826404PI 117060
209PI 116873No. 439Ilex sp.Hunan Sheng, China Historic1936COLLECTED10/1935Ma-Ling-Tung5641825132PI 116873
210PI 114757No. 100Ilex corallina Franch. China Historic1936COLLECTED1813205PI 114757
211PI 114758No. 101Ilex dunniana H. Lév. China Historic1936COLLECTED1813206PI 114758
212PI 114759No. 102Ilex pernyi Franch. China Historic1936COLLECTED1813207PI 114759
213PI 111359No. 18Ilex micrococca Maxim. Hubei Sheng, China Historic1935COLLECTEDWestern Hubei1803744PI 111359
214PI 111360No. 74Ilex micrococca Maxim. Hubei Sheng, China Historic1935COLLECTEDWestern Hubei1803745PI 111360
215PI 111388No. 21Ilex purpurea Hassk. Hubei Sheng, China Historic1935COLLECTEDWestern Hubei1803772PI 111388
216PI 110867Ilex cornuta Lindl. & Paxton California, United States Historic1935COLLECTED1492168PI 110867
217PI 110289Ilex pubescens Hook. & Arn. Guangdong Sheng, China Historic1935COLLECTED11/1934Chinchow on the North River3011801738PI 110289
218PI 108761Ilex aquifolium L. Bosnia and Herzegovina Historic1935COLLECTEDAbove Sarajevo1798990PI 108761
219PI 109296No. T-337Ilex sp.Bolu, Turkey Historic1935COLLECTED10/09/1934About 14.4km west of Bolu1568213PI 109296
220PI 107725Ilex opaca Aiton Unknown Historic1934COLLECTEDYugoslavia1797459PI 107725
221PI 107436'Argentea marginata'Ilex aquifolium L. New York, United States Historic1934COLLECTED1492678PI 107436
222PI 107437'Argentea medio-picta'Ilex aquifolium L. New York, United States Historic1934COLLECTED1796805PI 107437
223PI 107439CamelliIfolia (seedling)Ilex ×altaclerensis (Loudon) Dallim. New York, United States Historic1934COLLECTED1796807PI 107439
224PI 107440'Ciliata major'Ilex aquifolium L. New York, United States Historic1934COLLECTED1796808PI 107440
225PI 107441'Cookii'Ilex aquifolium L. New York, United States Historic1934COLLECTED1796809PI 107441
226PI 107442'Flavescens'Ilex aquifolium L. New York, United States Historic1934COLLECTED1796810PI 107442
227PI 107444'Integrifolia'Ilex aquifolium L. New York, United States Historic1934COLLECTED1796812PI 107444
228PI 107445'Rubicaulis aurea'Ilex aquifolium L. New York, United States Historic1934COLLECTED1796813PI 107445
229PI 107446Ilex sp.New York, United States Historic1934COLLECTED1796814PI 107446
230PI 107447Ilex sp.New York, United States Historic1934COLLECTED1796815PI 107447
231PI 107448Ilex perado Aiton subsp. platyphylla (Webb & Berthel.) S. Andrews New York, United States Historic1934COLLECTED1796816PI 107448
232PI 107626Ilex aquifolium L. Bosnia and Herzegovina Historic1934COLLECTEDSarajevo1797032PI 107626
233PI 105888Ilex sp.Uttar Pradesh, India Historic1934COLLECTEDSarahan1793810PI 105888
234PI 105610No. F. 87Ilex sp.Jiangsu Sheng, China Historic1934COLLECTED09/15/1933Near Ta Tseh Tsuen, Yung Hsien3011794445PI 105610
235PI 112222No. 47Ilex sp. China Historic1934COLLECTEDTa Tseh Tsuen, Yung Hsien1491957PI 112222
236PI 105320Ilex purpurea Hassk. China Historic1934COLLECTED1793948PI 105320
237PI 105321Ilex rotunda Thunb. China Historic1934COLLECTED1793949PI 105321
238PI 105322Ilex viridis Champ. ex Benth. China Historic1934COLLECTED1793950PI 105322
239PI 105323Ilex wilsonii Loes. China Historic1934COLLECTED1793951PI 105323
240PI 103698Ilex paraguariensis A. St.-Hil. Paraguay Historic1933COLLECTED1493572PI 103698
241PI 103447No. 24323Ilex sp.Sichuan Sheng, China Historic1933COLLECTEDMutirong, Muli Territory33111791781PI 103447
242PI 103459No. 22299Ilex sp. China Historic1933COLLECTEDSolola, Tsarung Province37631791795PI 103459
243PI 103460No. 23654Ilex sp. China Historic1933COLLECTEDMountains of Kaakerbo, Tsarung Province30101791796PI 103460
244PI 103427No. D-335Ilex corallina Franch. China Historic1933COLLECTED1791763PI 103427
245PI 103221No. 24556Ilex pernyi Franch. Sichuan Sheng, China Historic1933COLLECTEDBetween Muli and Kulu27091791523PI 103221
246PI 102905Ilex hookeri King India Historic1933COLLECTED1791234PI 102905
247PI 102906Ilex insignis Hook. f. India Historic1933COLLECTED1791235PI 102906
248PI 102944No. 5Ilex sp.Jiangxi Sheng, China Historic1933COLLECTEDHuangyensze1791273PI 102944
249PI 102945No. 53Ilex sp.Jiangxi Sheng, China Historic1933COLLECTEDKuanyinchiao1791274PI 102945
250PI 102946No. 68Ilex sp.Jiangxi Sheng, China Historic1933COLLECTEDPailohtung1791275PI 102946
251PI 105222Ilex paraguariensis A. St.-Hil. Argentina Historic1933COLLECTED1492422PI 105222
252PI 102299Ilex pedunculosa Miq. China Historic1933COLLECTEDManchuria1790478PI 102299
253PI 102300Ilex pedunculosa Miq. China Historic1933COLLECTEDManchuria1790480PI 102300
254PI 102301Ilex pedunculosa Miq. China Historic1933COLLECTEDManchuria1790481PI 102301
255PI 102075Ilex purpurea Hassk. China Historic1933COLLECTED1789903PI 102075
256PI 101516Ilex paraguariensis A. St.-Hil. Argentina Historic1932COLLECTED1789025PI 101516
257PI 100561F. 29908Ilex micrococca Maxim. China Historic1932COLLECTED1788166PI 100561
258PI 100562F. 30561Ilex sp. China Historic1932COLLECTED1788167PI 100562
259PI 99876F. 30479Ilex venulosa Hook. f. China Historic1932COLLECTED1787192PI 99876
260PI 99167Pa Hua MuIlex micrococca Maxim. Guizhou Sheng, China Historic1932COLLECTED10/20/1931Forest slope near Machaoho, Fangchingshan, Kiangkouhsien10541786461PI 99167
261PI 99180No. 48Ilex sp.Guizhou Sheng, China Historic1932COLLECTED10/25/1931Roadside near Hweihsiangping15951786474PI 99180
262PI 99181No. 93Ilex sp.Guizhou Sheng, China Historic1932COLLECTED11/14/1931Tahoyen, Kiangkouhsien7531786475PI 99181
263PI 99236No. 55Ilex purpurea Hassk. Guizhou Sheng, China Historic1932COLLECTED10/25/1931Rocky slopes near Soochiapo, Fangchingshan, Kiangkouhsien9031786530PI 99236
264PI 97995'Pyramidalis'Ilex aquifolium L. Netherlands Historic1932COLLECTED1784625PI 97995
265PI 94775Ilex paraguariensis A. St.-Hil. Argentina Historic1931COLLECTED1781476PI 94775
266PI 94152No. 7968Ilex integra Thunb. Tôkyô, Japan Historic1931COLLECTED1781241PI 94152
267PI 94004No. 29054Ilex delavayi Franch. China Historic1931COLLECTED1781084PI 94004
268PI 93898No. 28997Ilex corallina Franch. China Historic1931COLLECTED1780970PI 93898
269PI 93899No. 29020Ilex yunnanensis Franch. China Historic1931COLLECTED1780971PI 93899
270PI 92775Ilex ×altaclerensis (Loudon) Dallim. France Historic1931COLLECTED1779529PI 92775
271PI 92776Ilex pernyi Franch. France Historic1931COLLECTED1779530PI 92776
272PI 92493Ilex integra Thunb. England, United Kingdom Historic1931COLLECTED1779337PI 92493
273PI 91960Ilex latifolia Thunb. Hyôgo, Japan Historic1931COLLECTED1778466PI 91960
274PI 91518Ilex latifolia Thunb. Kanagawa, Japan Historic1931COLLECTED1493915PI 91518
275PI 91252Ilex pedunculosa Miq. Hyôgo, Japan Historic1931COLLECTED1776062PI 91252
276PI 91253Ilex integra Thunb. Hyôgo, Japan Historic1931COLLECTED1776063PI 91253
277PI 91254Ilex purpurea Hassk. Hyôgo, Japan Historic1931COLLECTED1776064PI 91254
278PI 91255Ilex sugerokii Maxim. Hyôgo, Japan Historic1931COLLECTED1776065PI 91255
279PI 90792Ilex macropoda Miq. Unknown Historic1930COLLECTEDChosen1775437PI 90792
280PI 88236Ilex argentina Lillo Tucumán, Argentina Historic1930COLLECTED1752000PI 88236
281PI 86677Ilex fragilis Hook. f. West Bengal, India Historic1930COLLECTED1750473PI 86677
282PI 86678Ilex wightiana Wall. ex Wight West Bengal, India Historic1930COLLECTED1750474PI 86678
283PI 85870No. 3904Ilex rotunda Thunb. Sizuoka, Japan Historic1930COLLECTED01/24/19301749473PI 85870
284PI 83795No. 3155Ilex latifolia Thunb. Tôkyô, Japan Historic1930COLLECTED12/13/19291747326PI 83795
285PI 85735No. 1450Ilex serrata Thunb. Akita, Japan Historic1929COLLECTED10/11/19291749362PI 85735
286PI 80095Ilex purpurea Hassk. Georgia, United States Historic1929COLLECTED1496271PI 80095
287PI 80363No. 17368Ilex sp.Sichuan Sheng, China Historic1929COLLECTED01/1929In spruce forests, southern slopes ot Mount Gibbah, Muli34621744209PI 80363
288PI 80083Ilex insignis Hook. f. West Bengal, India Historic1929COLLECTED1743973PI 80083
289PI 80403Ilex pernyi Franch. Île-de-France, France Historic1929COLLECTED1744245PI 80403
290PI 78144NA 82707Ilex ciliospinosa Loes. Grand-Est, FranceNA1928COLLECTED1733747PI 78144
291PI 78145Ilex bioritsensis Hayata Grand-Est, France Historic1928COLLECTED1733748PI 78145
292PI 78146Ilex wilsonii Loes. Grand-Est, France Historic1928COLLECTED1733749PI 78146
293PI 77832No. 892-3Ilex serrata Thunb. Massachusetts, United States Historic1928COLLECTED1733382PI 77832
294PI 77583 NA 57'National'Ilex cornuta Lindl. & Paxton District of Columbia, United StatesNANot Available1928COLLECTEDCultivarCuttings taken from plant in National Botanic Garden by Paul Russell 1928--this plant is now called Ilex cornuta 'National' (From PI book: "From Washington, D.C. Plants grown from cuttings collected by Paul Russell, Bureau of Plant Industry. Received September 5, 1928. A form of the Chinese holly with nearly spineless leaves. These cuttings were taken from a pistillate shrub about 9 feet high, growing in the National Botanic Garden." See also NA 7516 for more info from Eisenbeiss regarding this plant's history. -SBL1495929PI 77583 NA 57
295PI 77165Ilex geniculata Maxim. Cienfuegos, Cuba Historic1928COLLECTED1732756PI 77165
296PI 76119No. 666Ilex pedunculosa Miq. Kyôto, Japan Historic1928COLLECTED02/1928Mountain side of Higashi-yama1497417PI 76119
297PI 75713SoyogoIlex sp.Kyôto, Japan Historic1927COLLECTED11/02/19271730404PI 75713
298PI 75716UmemodokiIlex serrata Thunb. Kyôto, Japan Historic1927COLLECTED11/12/1927Old garden ath Golden Pavilion, Kyoto1730410PI 75716
299PI 73278Ilex aquifolium L. Île-de-France, France Historic1927COLLECTED1728449PI 73278
300PI 73438Ilex yunnanensis Franch. England, United Kingdom Historic1927COLLECTED1728510PI 73438
301PI 70980Hung tung tsingIlex cornuta Lindl. & Paxton Jiangxi Sheng, China Historic1927COLLECTED12/1926En route from Kian to Taihop1725641PI 70980
302PI 70981Lak kokIlex sp.Jiangxi Sheng, China Historic1927COLLECTED12/1926En route from Kian to Wanan1725642PI 70981
303PI 70982Hung tung tsingIlex sp.Jiangxi Sheng, China Historic1927COLLECTED12/1926En route from Kian to Wanan1725643PI 70982
304PI 70983Laap shueIlex sp.Jiangxi Sheng, China Historic1927COLLECTED12/16/1926Near Kanchow1725644PI 70983
305PI 70979Lak kokIlex cornuta Lindl. & Paxton Jiangxi Sheng, China Historic1927COLLECTED12/08/1926Near Kian1116163PI 70979
306PI 71859No. 931Ilex canariensis Poir. Canarias, Spain Historic1927COLLECTED12/23/1926Orotava Botanic Gardens, Teneriffe1726617PI 71859
307PI 72467No. 6925Ilex sp. Myanmar Historic1927COLLECTED1727329PI 72467
308PI 69865Ilex paraguariensis A. St.-Hil. Asunción, Paraguay Historic1926COLLECTED1499661PI 69865
309PI 67361Ilex integra Thunb. England, United Kingdom Historic1926COLLECTED1719685PI 67361
310PI 67362Ilex latifolia Thunb. England, United Kingdom Historic1926COLLECTED1719686PI 67362
311PI 67363Ilex sikkimensis Kurz England, United Kingdom Historic1926COLLECTED1719687PI 67363
312PI 67032Ilex pernyi Franch. Île-de-France, France Historic1926COLLECTED1719405PI 67032
313PI 66028Ilex purpurea Hassk. Jiangsu Sheng, China Historic1926COLLECTED1499087PI 66028
314PI 65256Ilex insignis Hook. f. West Bengal, India Historic1925COLLECTED1716527PI 65256
315PI 64255No. 6249Ilex sp.Xizang Zizhiqu, China Historic1925COLLECTED1665572PI 64255
316PI 63793Chue t'in shue, Tsau peng shueIlex rotunda Thunb. Guangdong Sheng, China Historic1925COLLECTED1665180PI 63793
317PI 63794Pak lan heungIlex sp.Guangdong Sheng, China Historic1925COLLECTED02/16/19251665181PI 63794
318PI 63626Pak lan heungIlex rotunda Thunb. Guangdong Sheng, China Historic1925COLLECTED01/02/19251664999PI 63626
319PI 62810Ilex sikkimensis Kurz West Bengal, India Historic1925COLLECTED1664091PI 62810
320PI 62723Ilex ciliospinosa Loes. England, United Kingdom Historic1925COLLECTED1664030PI 62723
321PI 60646Ilex insignis Hook. f. West Bengal, India Historic1924COLLECTED1662251PI 60646
322PI 59391Ilex latifolia Thunb. England, United Kingdom Historic1924COLLECTED1660681PI 59391
323PI 57770Ilex paraguariensis A. St.-Hil. Buenos Aires, Argentina Historic1923COLLECTED1502813PI 57770
324PI 57085Ilex sp.Hubei Sheng, China Historic1923COLLECTED1657455PI 57085
325PI 57086Ilex sp.Hubei Sheng, China Historic1923COLLECTED1657456PI 57086
326PI 65860Ilex cornuta Lindl. & Paxton Hubei Sheng, China Historic1923COLLECTED1499070PI 65860
327PI 56314No. 6781Ilex sp. China Historic1923COLLECTED10/192236001656219PI 56314
328PI 66866Ilex intricata Hook. f. West Bengal, India Historic1923COLLECTEDDarjilling1499253PI 66866
329PI 55621Ilex paraguariensis A. St.-Hil. Buenos Aires, Argentina Historic1922COLLECTED1502278PI 55621
330PI 55682Ilex insignis Hook. f. West Bengal, India Historic1922COLLECTED1654984PI 55682
331PI 55566Ilex paraguariensis A. St.-Hil. Asunción, Paraguay Historic1922COLLECTED1502240PI 55566
332PI 55493Ilex paraguariensis A. St.-Hil. Île-de-France, France Historic1922COLLECTED1502204PI 55493
333PI 55489Ilex paraguariensis A. St.-Hil. Buenos Aires, Argentina Historic1922COLLECTED1502201PI 55489
334PI 53452Ilex sp.Uttar Pradesh, India Historic1921COLLECTED1500573PI 53452
335PI 52644Ilex fargesii Franch. England, United Kingdom Historic1921COLLECTED1624556PI 52644
336PI 50420No. 1201Ilex sp.Massachusetts, United States Historic1920COLLECTED1618187PI 50420
337PI 47697Ilex fragilis Hook. f. West Bengal, India Historic1919COLLECTED1613255PI 47697
338PI 47698Ilex insignis Hook. f. West Bengal, India Historic1919COLLECTED1613249PI 47698
339PI 47699Ilex intricata Hook. f. West Bengal, India Historic1919COLLECTED1613250PI 47699
340PI 47309Ilex paraguariensis A. St.-Hil. Asunción, Paraguay Historic1919COLLECTED1508606PI 47309
341PI 46891Ilex paraguariensis A. St.-Hil. Al Qāhirah, Egypt Historic1918COLLECTED1508320PI 46891
342PI 46564Ilex paraguariensis A. St.-Hil. Al Qāhirah, Egypt Historic1918COLLECTED1508217PI 46564
343PI 44676'Alba de Llamas'Ilex paraguariensis A. St.-Hil. Salta, Argentina Historic1917COLLECTED1506782PI 44676
344PI 44335Ilex latifolia Thunb. Toscana, Italy Historic1917COLLECTED1506700PI 44335
345PI 43598Ilex paraguariensis A. St.-Hil. Concepción, Paraguay Historic1916COLLECTED1506377PI 43598
346PI 43456Ilex paraguariensis A. St.-Hil. Argentina Historic1916COLLECTED1608180PI 43456
347PI 39667Ilex fragilis Hook. f. West Bengal, India Historic1914COLLECTED1604339PI 39667
348PI 39668Ilex intricata Hook. f. West Bengal, India Historic1914COLLECTED1604340PI 39668
349PI 38858Ilex paraguariensis A. St.-Hil. Asunción, Paraguay Historic1914COLLECTED1518227PI 38858
350PI 34836Ilex corallina Franch. Centre-Val de Loire, France Historic1913COLLECTED1515010PI 34836
351PI 34655Ilex paraguariensis A. St.-Hil. Buenos Aires, Argentina Historic1912COLLECTED1514936PI 34655
352PI 34525Ilex fargesii Franch. Massachusetts, United States Historic1912COLLECTED05/19071599886PI 34525
353PI 34527Ilex pernyi Franch. Massachusetts, United States Historic1912COLLECTED1599888PI 34527
354PI 34546Ilex macrocarpa Oliv. Massachusetts, United States Historic1912COLLECTED19071599908PI 34546
355PI 34152Ilex paraguariensis A. St.-Hil. Asunción, Paraguay Historic1912COLLECTED1521166PI 34152
356PI 32945Ilex cornuta Lindl. & Paxton Liguria, Italy Historic1912COLLECTED1598546PI 32945
357PI 31255Ilex cornuta Lindl. & Paxton Liguria, Italy Historic1911COLLECTED1597068PI 31255
358PI 29134Ilex paraguariensis A. St.-Hil. Paraguay Historic1910COLLECTED1518190PI 29134
359PI 29097YerbaIlex paraguariensis A. St.-Hil. Unknown Historic1910COLLECTED1518164PI 29097
360PI 25529'Yerba'Ilex paraguariensis A. St.-Hil. Paraguay Historic1909COLLECTED1587524PI 25529
361PI 24638Chinese hollyIlex cornuta Lindl. & Paxton Shanghai Shi, China FRUITHistoric1909COLLECTED1522826PI 24638
362PI 24313Ilex paraguariensis A. St.-Hil. Buenos Aires, Argentina Historic1908COLLECTED1522683PI 24313
363PI 22979Tu hu tseIlex cornuta Lindl. & Paxton Jiangsu Sheng, China Historic1908COLLECTED04/26/19081585031PI 22979
364PI 19105Yerba mateIlex paraguariensis A. St.-Hil. Buenos Aires, Argentina Historic1906COLLECTED1532348PI 19105
365PI 10449Ilex crenata Thunb. Kanagawa, Japan Historic1904COLLECTED1524776PI 10449
366PI 8953Ilex paraguariensis A. St.-Hil. Buenos Aires, Argentina Historic1902COLLECTED1548251PI 8953
367PI 8664Ilex wightiana Wall. ex Wight Tamil Nadu, India Historic1902COLLECTED1547869PI 8664
368PI 7554Ilex integra Thunb. Île-de-France, France Historic1901COLLECTED1545978PI 7554
369PI 4594Ilex decidua Walter Missouri, United States Historic1900COLLECTED1539981PI 4594
370PI 4017Ilex verticillata (L.) A. Gray North Carolina, United States Historic1899COLLECTED1539121PI 4017
371PI 3035Ilex paraguariensis A. St.-Hil. France Historic1899COLLECTED1537220PI 3035
372PI 2173Ilex integra Thunb. France Historic1899COLLECTED1535863PI 2173
373PI 1698'Ilex balearica'Ilex aquifolium L. France Historic1898COLLECTED1534738PI 1698
374NA 89317NA 89317Ilex buergeri Miq. NANot Available2026Uncertain improvement statusAn almost certainly planted individual of this species was found growing on the northern side of the drainage/ravine in the woods just north of the Holly and Magnolia Collection. It may represent one of numerous accessions of this species that were brought to USNA in the past, and may have been "hidden" in this location by former staff.2197399NA 89317
375NA 89318NA 89318Ilex triflora Blume var. kanehirae (Yamam.) S. Y. Hu NANot Available2026Uncertain improvement statusPlant found growing on the northern side of the drainage/ravine in the woods just north of the Holly and Magnolia Collection. It may have been planted here, or could possibly have occurred as a spontaneous seedling from the collection, although this is the only such individual of this species to be found under such conditions and there are other plants nearby that suggest intentional planting.2197400NA 89318
376NA 89312(hybrid selection)Ilex hybr.NANot Available2026CloneTall, relatively upright-growing plant of clearly hybrid background (of the kind generally only produced at USNA in the past) found growing next to the old abandoned cold frame in the woods west of the fir collection. Based on its appearance, it could be 'Scepter' or it could be from the cross that produced 'Scepter', or a similar mix of species; the plant is female but only has some fruit on one side, probably due to excessive shade.2197394NA 89312
377NA 89313NA 89313Ilex aquifolium L. NANot Available2026Uncertain improvement statusLarge, but sparse and partly sprawling, layering plant (due to shade) found growing next to the abandoned cold frame in the woods west of the fir collection. The plant is very healthy and may be unusually well adapted to hot summers, although the conditions where the plant was found might be more favorable. It almost certainly had an accession number, and would not have arisen spontaneously, but no clear information about this cold frame or the plants around it has been located to date.2197395NA 89313
378NA 89314Senjoensis GroupIlex pedunculosa Miq. NANot Available2026Uncertain improvement statusFull name as recorded at the U.S. National Arboretum: Ilex pedunculosa Senjoensis Group. Large, multistemmed (stems widely spaced from below ground) plant, with a large portion seemingly prostrate and layering, found growing next to the abandoned cold frame in the woods west of the fir collection. The plant is unusually healthy for a member of this species in this area, although the conditions here might be especially favorable for hollies. It almost certainly had an accession number, and would not have arisen spontaneously, but no clear information about this cold frame or the plants around it has been located to date. Based on the presence of extensive prostrate growth, this is very likely to be either NA 51591 or 58525 (both Yinger 798, received from Brookside Gardens and originally collected in Japan), received as a prostrate form and updated here to I. pedunculosa var. senjoensis (no longer recognized as taxonomically distinct by most authorities) by T. R. Dudley and Gene Eisenbeiss. However, it is possible that some other accession that was collected from a high altitude, such as NA 37415, might have also been referable to Senjoensis Group.2197396NA 89314
379Ames 36712JDC/IL/2026/002/1166Ilex decidua Walter Missouri, United StatesNC7SEED2026COLLECTED02/04/2026South, southeast of Vienna approximately 2.65 miles just off highway 63 immediately east of intersection with county road 28, T39N R9W NE 1/4 Sec. 5 and also from NE 1/4 Sec. 16.38.15077900, -91.92982800259Roadside. Fenceline.Wild materialThe native range of Ilex decidua is concentrated throughout the southeastern United States as far west as central Texas and as far east as Virginia and as far north as northern Missouri (Little, E. L. Jr. 1977). Ilex decidua in its northern range of Missouri and Illinois seems to be rather restricted to the Mississippi and Illinois watersheds (Azotea, C., personal communication, 10 April 2026). Possumhaw is typically a densely, multi-stemmed shrub that can be trained into a tree potentially reaching 20' in height. White flowers are extremely small, inconspicuous. The species is dioecious (requiring male representation for fruit set), but some plants have been reported with perfect flowers. This species is known to inhabit limestone glades and bluffs, upland areas frequently along fence-lines and powerline cuts, but also in bottomland areas including sloughs and swamps. Specimens in upland habitats tend to be smaller statured specimens and seem to have smaller fruit loads (Azotea, C., personal communication, 10 April 2026). Ilex decidua has great adaptability in the urban landscape, with no major pests and great winter interest (Azotea, C., personal communication, 10 April 2026). Fruits on some genotypes of this species have been known to be orangish or even pink, but majority are red and typically persist well into late-winter. Persistence of fruit seems to vary across genotypes, where fruits on some genotypes dry/shrivel early and therefore drop early in winter, while others persist well into spring (Azotea, C., personal communication, 10 April 2026).  Fruit persistence can also be impacted by weather conditions and bird predation. This accession represents genetics from south-central Missouri growing in an upland habitat with red fruits measuring 6-8 mm in diameter with circa 3-4 seeds per fruit.  2193481Ames 36712
380NA 89199IleAno-HA-HNK-JIlex anomala Hook. & Arn. Hawaii, United StatesNANot Available2026COLLECTED07/14/2025Hakalau Forest National Wildlife Refuge: Middle Honokua, Hawai'i [bulk collection from an area spanning 19.340 deg. to 19.372 deg., and -155.889 deg. to -155.787 deg.; coordinates of record are approximately at the center of this area.]19.35600000, -155.83850000950Elevation: ca 800 to 1100 m.Wild material2193510NA 89199
381NA 89198IleAno-HA-HOO-GIlex anomala Hook. & Arn. Hawaii, United StatesNANot Available2025COLLECTED04/08/2025Hakalau Forest National Wildlife Refuge: Kona Unit: middle access road, Ho'okena, Hawai'i [bulk collection from an area spanning 19.370 deg. to 19.393 deg., and -155.858 deg. to -155.783 deg.; coordinates of record are approximately at the center of this area.]19.38150000, -155.82050000850Elevation: ca 700 to 1000 m.Wild material2193509NA 89198
382NA 89152'Bill Cannon'Ilex aquifolium L. Massachusetts, United StatesNANot Available2025DEVELOPEDPRE 2025CultivarFull name as recorded at the U.S. National Arboretum: Ilex aquifolium 'Bill Cannon'. Holly Society of America's Test Holly for 2026. According to the originator, this might be I. x meserveae, although it closely resembles I. aquifolium (J. Resch considers that it might be a backcross to I. aquifolium, if not pure I. aquifolium as it has been distributed by HSA).2192987NA 89152
383NA 89192TRx251105-01MDIlex laevigata (Pursh) A. Gray Maryland, United StatesNA2025COLLECTED11/05/2025Lake Waterford Park: south side of Lake Waterford.39.11352463, -76.56110440Plants were collected from the south side of Lake Waterford (a 12-acre lake fed by the Magothy River), where they were growing within 2 m of the water's edge.Wild material2193014NA 89192
384NA 891552025RJLSEUS-02Ilex longipes Chapm. ex Trel. Alabama, United StatesNA2025COLLECTED09/30/2025Scarham Creek: southeast of the Pea Ridge Road bridge, approximately 50-80 meters on the south side of the creek.34.32520700, -86.15967000261Plants are growing on relatively steep woodland slopes in shade. The woodlands are primarily deciduous with a few evergreens infrequently mixed. Slope: 15 to 30 percent. Aspect: Northeast. Soil: The primary soil type in the area of this collection is classified as Rockland Series sandstone (Rc). The Rockland series consists of well drained soils formed in loamy colluvium from rotational landslides on slopes of stream valleys and dissections of ground moraines. Saturated hydraulic conductivity is moderate in the upper part of the profile and moderately slow in the lower part. Slopes range from 18 to 70 percent. Some pedons have buried leaves, twigs, and moss. Mass soil movement has left random remnants of soil structure from prior upslope soil development and has buried tree branches, leaves and moss. The upper 0-2 cm of soil contains slightly decomposed organic matter while lower soils layers to a depth of 58 cm are comprised of dark reddish-brown to brown silt loam and are moderately acidic. Below 58 cm the soils are generally dark reddish-brown silt loam and strongly to moderately alkaline. (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions). Geology: The underlying geology of this area is of Pennsylvanian origin and characterized as Pottsville Formation. It is characterized by light-gray thin to thick-bedded quartzose sandstone and conglomerate containing interbedded dark-gray shale, siltstone, and coal. Mapped on Lookout Mountain, Blount and Chandler Mountains, and Sand Mountain northeast of Blount County, and on the mountains of Jackson, Marshall, and Madison Counties north and west of the TN river. (Source: USGS Alabama Geologic Map Data).Wild material2193001NA 89155
385NA 891602025RJLSEUS-08Ilex hybr.Florida, United StatesNA2025COLLECTED10/02/2025Eglin Air Force Base: coastal dune ridge on Eglin AFB behind businesses located along Florida State Highway 98. The site is accessed through a now gated and permanently closed Eglin AFB, East River Unit, A-21 Road (now open to pedestrian traffic only with a valid Eglin user permit). After walking through the gated road, the site is to the west along 4-wheeling trails.30.41191000, -86.773158008Part shade in a primarily evergreen extremely dry dune community with a mixed understory of deciduous and evergreen plants. Slope: 0 to 2 percent. Soils: The soils of this are characterized as Kureb Series sand, 0 to 8 percent slopes. The Kureb series consists of very deep, excessively drained, gently sloping to moderately steep soils on Coastal Plain uplands and on side slopes along streams and bays. They have formed in marine, aeolian, or fluvial sands. Slopes range from 0 to 20 percent. The thickness of the sandy horizon is more than 2 meters. Soil reaction is neutral to extremely acid throughout. All horizons are fine sand, sand, or coarse sand. Silt plus clay content is less than 5 percent. (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions). Geology: The underlying geology of this area is characterized as undifferentiated Quaternary Sediments. Much of Florida's surface is covered by a varying thickness of undifferentiated sediments consisting of siliciclastics, organics and freshwater carbonates. Where these sediments exceed 6.1 meters thick, they were mapped as discrete units. To subdivide the undifferentiated sediments, those sediments occurring in flood plains were mapped as alluvial and flood plain deposits (Qal). Sediments showing surficial expression of beach ridges and dunes were mapped separately (Qbd) as were the sediments composing Trail Ridge (Qtr). Terrace sands were not mapped (refer to Healy [1975] for a discussion of the terraces in Florida). The subdivisions of the Undifferentiated Quaternary Sediments (Qu) are not lithostratigraphic units but are utilized to facilitate a better understanding of the State's geology. The siliciclastics are light gray, tan, brown to black, unconsolidated to poorly consolidated, clean to clayey, silty, unfossiliferous, variably organic-bearing sands to blue green to olive green, poorly to moderately consolidated, sandy, silty clays. Gravel is occasionally present in the panhandle. Organics occur as plant debris, roots, disseminated organic matrix and beds of peat. Freshwater carbonates, often referred to as marls in the literature, are scattered over much of the State. In southern Florida, freshwater carbonates are nearly ubiquitous in the Everglades. These sediments are buff colored to tan, unconsolidated to poorly consolidated, fossiliferous carbonate muds. Sand, silt and clay may be present in limited quantities. These carbonates often contain organics. The dominant fossils in the freshwater carbonates are mollusks. (Source: USGS Florida Geologic Map Data).Wild material2193006NA 89160
386NA 89181WV25-01Ilex montana Torr. & A. Gray West Virginia, United StatesNA2025COLLECTED09/12/2025Monongahela National Forest: found while driving down US-250, on the south side of the road, beyond the drainage cut.38.60535544, -79.861143411127Wild materialHerbarium voucher deposited USNA2192760NA 89181
387NA 89184WV25-05Ilex collina Alexander West Virginia, United StatesNA2025COLLECTED09/12/2025Monongahela National Forest. The only individual found during a 0.6-mile transect on the SW side of US 250.38.60075654, -79.854920121134A relatively open, sunny moor with thin soil.Wild materialHerbarium voucher deposited USNA2192764NA 89184
388NA 89185WV25-06Ilex collina Alexander West Virginia, United StatesNA2025COLLECTED09/12/2025Monongahela National Forest. This population was on the NE side of US-250.38.60211468, -79.854262121124Growing in mostly shade within a densely vegetated peaty bog.Wild materialHerbarium voucher deposited USNA2192765NA 89185
389NA 89186WV25-07Ilex collina Alexander West Virginia, United StatesNA2025COLLECTED09/12/2025Manongahela National Forest.38.61324900, -79.868596521100Plants occurring on a north facing slope at an intersection where they were routinely cut back, with resulting runners and ramets.Wild materialHerbarium voucher deposited USNA2192766NA 89186
390NA 89188WV25-10Ilex collina Alexander West Virginia, United StatesNA2025COLLECTED09/13/2025Monongahela National Forest.38.61750465, -79.874007241110South-facing slope in sunny conditions along forest edge; peaty roadside ditch at the edge of spruce forest.Wild materialHerbarium voucher deposited USNA2192768NA 89188
391NA 89190WV25-12Ilex collina Alexander West Virginia, United StatesNA2025COLLECTED09/13/2025Monongahela National Forest.38.52413054, -79.937892231136Steep, southwest-facing slope above parking area.Wild materialHerbarium voucher deposited USNA2192770NA 89190
392NA 89191WV25-14Ilex montana Torr. & A. Gray West Virginia, United StatesNA2025COLLECTED09/13/2025Monongahela National Forest: east side of Forest Road 233.38.57289483, -79.900668431109Growing on east side of road.Wild materialHerbarium voucher deposited USNA2192772NA 89191
393NA 89139(undetermined cultivar)Ilex opaca Aiton NANot Available2025Uncertain improvement statusMature male Ilex opaca growing in the center of the Lee Garden (bed AZ-LG-CTR) just below the pond in the vicinity of accessioned female cultivars/clones, and appearing to be similar in age, so presumably planted with them as a pollinator. -SBL2192305NA 89139
394NA 89129TRx250809-01PAIlex mucronata (L.) M. Powell et al. Pennsylvania, United StatesNA2025COLLECTED07/09/2025Hickory Run Boulder Field41.04973900, -75.64300700555Gowing within ca 15 m of boulder field edge, most individuals occured on sun-exposed margin.Wild material2191815NA 89129
395NA 89112NA 89112Ilex crenata Thunb. NANot Available2025Uncertain improvement statusOne plant long considered and mapped as an individual of NA 44889, Ilex crenata var. paludosa (at least as early as 1993) is very different in habit from the lower, spreading plant of NA 44889 next to it. It is unclear if this could have been another accession, or if seed collected from var. paludosa could have outcrossed with another taxon to result in this growth habit. Assigned a new number out of an abundance of caution; likely to have been collected from the wild in Japan in any case, since all of the plants in the bed AC-J-L planting were.The plant might be male or non-flowering in 2025, as there are no fruit present, and I did not see any floral remants.2191658NA 89112
396NA 88915TRx250318-01FLIlex vomitoria Aiton Florida, United StatesNANot Available2025COLLECTED03/18/2025Yellow River Wildlife Management Area: along dirt access road DOF Y15, in the vicinity fo Burnt Grocery Creek.30.62922000, -86.84158000Understory of dry pine forest.Wild materialsmall leaved plant2172685NA 88915
397NA 88747TXWL2025-038Ilex opaca Aiton Texas, United StatesNA2025COLLECTED01/23/2025Milam, Texas: On flat road, private land.31.43288000, -93.88208000Upland shortleaf pine stand with sandy loam; closed canopy.Wild material2171021NA 88747
398NA 88746TXWL2025-039Ilex verticillata (L.) A. Gray Louisiana, United StatesNA2025COLLECTED01/24/2025Kistachie National Forest: Compartment 49, Indian Creek.31.09110000, -93.53333000Slough/baygall area with standing water below a longleaf pine stand.Wild material2171020NA 88746
399NA 88981TXWL2025-031Ilex opaca Aiton Texas, United StatesNA2025COLLECTED01/07/2025South of Jericho community.31.67015000, -94.25641000Western sandy lands; small grove of Ilex opaca in upland position above a baygall. Bare ground cover with Ilex opaca regeneration. Adjacent to a loblolly pine plantation and a new longleaf pine plantation.Wild material2170965NA 88981
400NA 88982TXWL2025-032Ilex opaca Aiton Texas, United StatesNA2025COLLECTED01/07/2025South of Jericho community.31.66973000, -94.25613000Western sandy lands; edge of small grove of Ilex opaca with sun exposure in upland position above a baygall. Bare ground cover with Ilex opaca regeneration. Adjacent to a new longleaf pine plantation located on fireline.Wild material2170966NA 88982
401NA 88983TXWL2025-033Ilex opaca Aiton Texas, United StatesNA2025COLLECTED01/12/2025Adjacent to Toledo Bend reservoir in Huxley Bay community, eastern Shelby County.31.77133000, -93.85094000Low area with heavy duff near reservoir.Wild material2170967NA 88983
402NA 88984TXWL2025-034Ilex opaca Aiton Texas, United StatesNA2025COLLECTED01/13/2025Fairmount Township: Sabine National Forest C-139: Foxhunters Hill.31.18153000, -93.72368000Baygall below a longleaf pine stand. Same system as 035 [NA 88985].Wild material2170968NA 88984
403NA 88985TXWL2025-035Ilex opaca Aiton Texas, United StatesNA2025COLLECTED01/13/2025Fairmount Township: Sabine National Forest C-139: Foxhunters Hill.31.18220000, -93.72291000Baygall below a longleaf pine stand. Same system as 034 [NA 88984].Wild material2170969NA 88985
404NA 88986TXWL2025-036Ilex opaca Aiton Texas, United StatesNANot Available2025COLLECTED01/13/2025Fairmount Township: Sabine National Forest C-139: Foxhunters Hill.31.18440000, -93.72253000Baygall below a longleaf pine stand.Wild material2170970NA 88986
405NA 88987TXWL2025-037Ilex opaca Aiton Texas, United StatesNA2025COLLECTED01/13/2025Fairmount Township: Sabine National Forest C-139: Foxhunters Hill.31.18365000, -93.72211000Baygall below a longleaf pine stand.Wild material2170971NA 88987
406NA 89095SNR2024Ilex decidua Walter Missouri, United StatesNA2024COLLECTED12/03/2024Shaw Nature Reserve: near Shaw Nature Reserve's Trail House.38.46697200, -90.81925000Upland site.Wild material2169943NA 89095
407NA 885772024RJLFL-001Ilex myrtifolia Walter Florida, United StatesNA2024COLLECTED11/12/2024Yellow River Marsh Preserve State Park: collected along trail from the parking lot. The property is accessed from a parking area located on Dickerson City Road.30.48472300, -87.071104005Mostly found in moist flatwoods in mixed evergreen and deciduous forest. Slope: N/A. Aspect: Various. The underlying geology of this area is characterized as undifferentiated Quaternary Sediments. Much of Florida's surface is covered by a varying thickness of undifferentiated sediments consisting of siliciclastics, organics and freshwater carbonates. Where these sediments exceed 6.1 meters thick, they were mapped as discrete units. In an effort to subdivide the undifferentiated sediments, those sediments occurring in flood plains were mapped as alluvial and flood plain deposits (Qal). Sediments showing surficial expression of beach ridges and dunes were mapped separately (Qbd) as were the sediments composing Trail Ridge (Qtr). Terrace sands were not mapped (refer to Healy [1975] for a discussion of the terraces in Florida). The subdivisions of the Undifferentiated Quaternary Sediments (Qu) are not lithostratigraphic units but are utilized in order to facilitate a better understanding of the State's geology. The siliciclastics are light gray, tan, brown to black, unconsolidated to poorly consolidated, clean to clayey, silty, unfossiliferous, variably organic-bearing sands to blue green to olive green, poorly to moderately consolidated, sandy, silty clays. Gravel is occasionally present in the panhandle. Organics occur as plant debris, roots, disseminated organic matrix and beds of peat. Freshwater carbonates, often referred to as marls in the literature, are scattered over much of the State. In southern Florida, freshwater carbonates are nearly ubiquitous in the Everglades. These sediments are buff colored to tan, unconsolidated to poorly consolidated, fossiliferous carbonate muds. Sand, silt and clay may be present in limited quantities. These carbonates often contain organics. The dominant fossils in the freshwater carbonates are mollusks. (Source: USGS Florida Geologic Map Data). The Goldhead Series consists of poorly drained and very poorly drained soils on broad areas of the flatwoods and in depressions. They formed in thick beds of stratified unconsolidated loamy and sandy marine sediments. Slopes range from 0 to 5 percent. The solum thickness ranges from 35 to greater than 60 inches. Soil reaction ranges from very strongly acid to mildly alkaline in the A and E horizons, and very strongly acid to moderately alkaline in the B and C horizons. (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions).Wild material2169914NA 88577
408NA 885782024RJLFL-004Ilex vomitoria Aiton Florida, United StatesNA2024COLLECTED11/12/2024Collected along Florida National Scenic Trail/Eglin Air Force Base unnamed road.30.44526100, -86.860947004Mixed evergreen and deciduous forests in part shade to full sun. Slope: 5 to 10 percent. Aspect: Various. The underlying geology of this area is characterized as Citronelle Formation of the Pliocene period. The Citronelle Formation is widespread in the Gulf Coastal Plain. The type section for the Citronelle Formation, named by Matson (1916), is near Citronelle, Alabama. The Citronelle Formation grades laterally, through a broad facies transition, into the Miccosukee Formation of the eastern Florida panhandle. Coe (1979) investigated the Citronelle Formation in portions of the western Florida panhandle. The Citronelle Formation is a siliciclastic, deltaic deposit that is lithologically similar to, and time equivalent with, the Cypresshead Formation and, at least in part, the Long Key Formation (Cunningham et al., 1998) of the peninsula. In the western panhandle, some of the sediments mapped as Citronelle Formation may be reworked Citronelle. The lithologies are the same and there are few fossils present to document a possible younger age. The Citronelle Formation consists of gray to orange, often mottled, unconsolidated to poorly consolidated, very fine to very coarse, poorly sorted, clean to clayey sands. It contains significant amounts of clay, silt and gravel which may occur as beds and lenses and may vary considerably over short distances. Limonite nodules and limonite-cemented beds are common. Marine fossils are rare but fossil pollen, plant remains and occasional vertebrates are found. Much of the Citronelle Formation is highly permeable. It forms the Sand and Gravel Aquifer of the surficial aquifer system contain organics. The dominant fossils in the freshwater carbonates are mollusks. (Source: USGS Florida Geologic Map Data). The soils of this are characterized as Lakeland Series Sand, 5 to 12 percent slopes. The Lakeland series consists of very deep, excessively drained, rapid to very rapidly permeable soils on uplands. They formed in thick beds of eolian or marine and/or fluvio-marine sands in the Southern Coastal Plain MLRA (133A), the Carolina and Georgia Sandhills (MLRA 137), the Eastern Gulf Coast Flatwoods (MLRA 152A) and the Atlantic Coast Flatwoods (MLRA 153A). Slopes are dominantly from 0 to 12 percent but can range to 85 percent in dissected areas. Thickness of the sand exceeds 200 cm. Silt plus clay in the 25 to 100 cm control section ranges from 5 to 10 percent. Reaction ranges from very strongly acid to moderately acid throughout. (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions).Wild material2169915NA 88578
409NA 885792024RJLFL-007Ilex myrtifolia Walter Florida, United StatesNA2024COLLECTED11/13/2024Wetlands accessed from Miller Bluff Road.30.62251200, -86.860307008Open, wet areas and woodland edge slightly above permanently wet habitat in part shade to full sun. Slope: 0 percent. Aspect: N/A.Wild material2169916NA 88579
410NA 885802024RJLFL-009Ilex myrtifolia Walter Florida, United StatesNA2024COLLECTED11/13/2024Woodlands on the north side of Fisher Old Mill Road about 0.10 kilometers west of intersection with DOF Y12.30.62693800, -86.839622006Mixed evergreen and deciduous moist to wet forest in part shade to full sun. From the associated taxa, this site can be quite wet during the growing season; however, it was dry during this collection time; likely due to extended drought. Slope: 0 percent. Aspect: South.Wild material2169917NA 88580
411NA 885812024RJLFL-012Ilex ambigua Torr. Florida, United StatesNA2024COLLECTED11/13/2024Woodland edge along the south side of Bud Bass Road (DOF B48), 0.30 kilometers east of the junction with Norman Riley Road.30.75763200, -86.8018430032Woodland edge. Slope: 3 percent. Aspect: North. The underlying geology of this area is characterized as Citronelle Formation of the Pliocene period. The Citronelle Formation is widespread in the Gulf Coastal Plain. The type section for the Citronelle Formation, named by Matson (1916), is near Citronelle, Alabama. The Citronelle Formation grades laterally, through a broad facies transition, into the Miccosukee Formation of the eastern Florida panhandle. Coe (1979) investigated the Citronelle Formation in portions of the western Florida panhandle. The Citronelle Formation is a siliciclastic, deltaic deposit that is lithologically similar to, and time equivalent with, the Cypresshead Formation and, at least in part, the Long Key Formation (Cunningham et al., 1998) of the peninsula. In the western panhandle, some of the sediments mapped as Citronelle Formation may be reworked Citronelle. The lithologies are the same and there are few fossils present to document a possible younger age. The Citronelle Formation consists of gray to orange, often mottled, unconsolidated to poorly consolidated, very fine to very coarse, poorly sorted, clean to clayey sands. It contains significant amounts of clay, silt and gravel which may occur as beds and lenses and may vary considerably over short distances. Limonite nodules and limonite-cemented beds are common. Marine fossils are rare but fossil pollen, plant remains and occasional vertebrates are found. Much of the Citronelle Formation is highly permeable. It forms the Sand and Gravel Aquifer of the surficial aquifer system contain organics. The dominant fossils in the freshwater carbonates are mollusks. (Source: USGS Florida Geologic Map Data). The soils of this are characterized as Lakeland Series Sand, 0 to 5 percent slopes. The Lakeland series consists of very deep, excessively drained, rapid to very rapidly permeable soils on uplands. They formed in thick beds of eolian or marine and/or fluvio-marine sands in the Southern Coastal Plain MLRA (133A), the Carolina and Georgia Sandhills (MLRA 137), the Eastern Gulf Coast Flatwoods (MLRA 152A) and the Atlantic Coast Flatwoods (MLRA 153A). Slopes are dominantly from 0 to 12 percent but can range to 85 percent in dissected areas. Thickness of the sand exceeds 200 cm. Silt plus clay in the 25 to 100 cm control section ranges from 5 to 10 percent. Reaction ranges from very strongly acid to moderately acid throughout. (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions).Wild material2169918NA 88581
412NA 885832024RJLFL-014Ilex amelanchier M. A. Curtis ex Chapm. Florida, United StatesNA2024COLLECTED11/13/2024Yellow River Water Management Area: found in an area where an unnamed unpaved road dead ends near the Yellow River. This unpaved road is accessed from Old River Road to the east near the junction with Shockley Springs Road. Individual plants for this collection were accessed by foot along the edge of the river and associated wetland depressions to the east of the road. The sampling for this portion of the population extends from the GPS coordinates listed above to the northeast where sampling ended at coordinates 30.871238 deg. N, 86.590170 deg. W.30.86989400, -86.5921850023Plants growing in seasonally inundated swamp habitats associated with the Yellow River and small tributaries. Plants are emergent during portions of the year, primarily during the growing season. During this time of collection, water had receded from the wetlands and the area was relatively dry. Slope: N/A. Aspect: N./A. The underlying geology of this area is the Alum Bluff Group of Miocene origin. It includes the Chipola Formation, Oak Grove Sand, Shoal River Formation, Choctawhatchee Formation and the Jackson Bluff Formation. The formations included in this group are generally defined based on their molluscan faunas and stratigraphic position. Puri (1953) described sediment facies as they relate to the formations of the Alum Bluff Group. These sediments are lithologically distinct as a group, not as individual units. The Alum Bluff Group crops out or is beneath a thin overburden in the western panhandle from river valleys in Okaloosa County eastward to western Jackson County. The Alum Bluff Group consists of clays, sands and shell beds which may vary from fossiliferous, sandy clays to unfossiliferous sands and clays and occasional carbonate beds. Mica is a common constituent and glauconite and phosphate occur sporadically. Induration varies from essentially nonindurated in sands to well indurated in carbonate lenses. Colors range from cream to olive gray with mottled reddish brown in weathered sections. Sand grain size varies from very fine to very coarse with sporadic occurrences of gravel. These sediments generally have low permeabilities and are part of the intermediate confining unit/aquifer system. (Source: USGS Florida Geologic Map Data). The surface soil profile of this area is primarily Kinston, Johnston, and Bibb soils, frequently flooded. Kinston Series soils are poorly drained, alluvial, fine-loamy, siliceous, and acid soils of flood plains. They are peaty to loamy friable, slightly sticky, and slightly plastic in the upper 0 to 7.5 cm of the strata with less sand and more silt-loam in progressively deeper strata. The Johnston Series consists of very deep, very poorly drained coarse-loamy, siliceous, active, acid, thermic Cumulic Humaquepts. The upper 0-75 cm is black, mucky loam that is friable and very strongly acid. The lower strata to 1.75 m are dark gray, loamy fine sand, single grained, loose and very strongly acid. (Source: NRCS Web Soil Survey).Wild material2169920NA 88583
413NA 885902024RM-003Ilex verticillata (L.) A. Gray Florida, United StatesNA2024COLLECTED11/21/2024Along the west/north side of the Yellow River about 0.33 kilometers (as the crow flies) upstream from the channel that leads the Fisher Old Mill Road Boat Ramp. This site was accessed by boat.30.62868600, -86.807480004Plants are found growing primarily in permanently saturated to submersed conditions along the edge of the Yellow River in part shade. The riverine habitat is comprised of a wide array of deciduous and evergreen trees, shrubs, and perennial plant species. Most of the plants along the river are adapted to acidic conditions and, while the aquatic habitat is generally fresh water, there is occasionally tidal or hurricane-caused saltwater intrusions. Slope: 0 percent. Aspect: East-southeast. The underlying geology of this area is characterized as Alluvium from undifferentiated Quaternary Sediments. Much of Florida's surface is covered by a varying thickness of undifferentiated sediments consisting of siliciclastics, organics and freshwater carbonates. Where these sediments exceed 20 feet (6.1 meters) thick, they were mapped as discrete units. To subdivide the undifferentiated sediments, those sediments occurring in flood plains were mapped as alluvial and flood plain deposits (Qal). Sediments showing surficial expression of beach ridges and dunes were mapped separately (Qbd) as were the sediments composing Trail Ridge (Qtr). Terrace sands were not mapped (refer to Healy [1975] for a discussion of the terraces in Florida). The subdivisions of the Undifferentiated Quaternary Sediments (Qu) are not lithostratigraphic units but are utilized to facilitate a better understanding of the State's geology. The siliciclastics are light gray, tan, brown to black, unconsolidated to poorly consolidated, clean to clayey, silty, unfossiliferous, variably organic-bearing sands to blue green to olive green, poorly to moderately consolidated, sandy, silty clays. Gravel is occasionally present in the panhandle. Organics occur as plant debris, roots, disseminated organic matrix and beds of peat. Freshwater carbonates, often referred to as marls in the literature, are scattered over much of the State. In southern Florida, freshwater carbonates are nearly ubiquitous in the Everglades. These sediments are buff colored to tan, unconsolidated to poorly consolidated, fossiliferous carbonate muds. Sand, silt and clay may be present in limited quantities. These carbonates often contain organics. The dominant fossils in the freshwater carbonates are mollusks. (Source: USGS Florida Geologic Map Data). The soils of this are characterized as Bibb-Kinston Series association. The Bibb Series consists of very deep, poorly drained, moderately permeable soils on flood plains of streams in the Southern Coastal Plain (133A) Major Land Resource Area. They formed in stratified loamy and sandy alluvium that are commonly and frequently flooded and water runs off the surface very slowly. Slopes range from 0 to 2 percent. Reaction ranges from extremely acid to strongly acid throughout. Content of mica flakes ranges from none to common. Content of rounded gravel typically ranges from 0 to 10 percent throughout, but may range to 50 percent in thin strata below a depth of 1 meter. Buried soil horizons, present in many pedons have the same range in color and texture as the surface horizon which has iron and organic matter concentrations in shades of brown and yellow ranging from none to common. Texture is sand, loamy sand, loamy fine sand, fine sandy loam, sandy loam, loam or silt loam. The Kinston Series consists of very deep, poorly drained, moderately permeable soils found on flood plains. They were formed in alluvial depositions. Slopes range from 0 to 2 percent. Soil reaction ranges from strongly acid to very strongly acid. Typical texture across the pedons is loamy, gravelly loam, sandy loam, and clay loam with dark concretions often present. (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions).Wild material2169927NA 88590
414NA 885912024RM-004Ilex opaca Aiton Florida, United StatesNA2024COLLECTED11/21/2024Along the west/north side of the Yellow River about 5.5 kilometers upstream from the State Highway 87 boat ramp. This site was accessed by boat.30.59986000, -86.879360002Plants are found growing primarily in permanently saturated to submersed conditions along the edge of the Yellow River in part shade. The riverine habitat is comprised of a wide array of deciduous and evergreen trees, shrubs, and perennial plant species. Most of the plants along the river are adapted to acidic conditions and, while the aquatic habitat is generally fresh water, there is occasionally tidal or hurricane-caused saltwater intrusions. Slope: 0 percent. Aspect: East-southeast. The underlying geology of this area is characterized as Alluvium from undifferentiated Quaternary Sediments. Much of Florida's surface is covered by a varying thickness of undifferentiated sediments consisting of siliciclastics, organics and freshwater carbonates. Where these sediments exceed 20 feet (6.1 meters) thick, they were mapped as discrete units. To subdivide the undifferentiated sediments, those sediments occurring in flood plains were mapped as alluvial and flood plain deposits (Qal). Sediments showing surficial expression of beach ridges and dunes were mapped separately (Qbd) as were the sediments composing Trail Ridge (Qtr). Terrace sands were not mapped (refer to Healy [1975] for a discussion of the terraces in Florida). The subdivisions of the Undifferentiated Quaternary Sediments (Qu) are not lithostratigraphic units but are utilized to facilitate a better understanding of the State's geology. The siliciclastics are light gray, tan, brown to black, unconsolidated to poorly consolidated, clean to clayey, silty, unfossiliferous, variably organic-bearing sands to blue green to olive green, poorly to moderately consolidated, sandy, silty clays. Gravel is occasionally present in the panhandle. Organics occur as plant debris, roots, disseminated organic matrix and beds of peat. Freshwater carbonates, often referred to as marls in the literature, are scattered over much of the State. In southern Florida, freshwater carbonates are nearly ubiquitous in the Everglades. These sediments are buff colored to tan, unconsolidated to poorly consolidated, fossiliferous carbonate muds. Sand, silt and clay may be present in limited quantities. These carbonates often contain organics. The dominant fossils in the freshwater carbonates are mollusks. (Source: USGS Florida Geologic Map Data). The soils of this are characterized as Bibb-Kinston Series association. The Bibb Series consists of very deep, poorly drained, moderately permeable soils on flood plains of streams in the Southern Coastal Plain (133A) Major Land Resource Area. They formed in stratified loamy and sandy alluvium that are commonly and frequently flooded and water runs off the surface very slowly. Slopes range from 0 to 2 percent. Reaction ranges from extremely acid to strongly acid throughout. Content of mica flakes ranges from none to common. Content of rounded gravel typically ranges from 0 to 10 percent throughout, but may range to 50 percent in thin strata below a depth of 1 meter. Buried soil horizons, present in many pedons have the same range in color and texture as the surface horizon which has iron and organic matter concentrations in shades of brown and yellow ranging from none to common. Texture is sand, loamy sand, loamy fine sand, fine sandy loam, sandy loam, loam or silt loam. The Kinston Series consists of very deep, poorly drained, moderately permeable soils found on flood plains. They were formed in alluvial depositions. Slopes range from 0 to 2 percent. Soil reaction ranges from strongly acid to very strongly acid. Typical texture across the pedons is loamy, gravelly loam, sandy loam, and clay loam with dark concretions often present. (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions).Wild material2169928NA 88591
415NA 885922024RM-005Ilex sp.Florida, United StatesNA2024COLLECTED11/21/2024Along the west/north side of the Yellow River about 1.1 kilometers upstream from the State Highway 87 boat ramp. This site was accessed by boat.30.57956000, -86.919164001Plants are found growing primarily in permanently saturated to submersed conditions along the edge of the Yellow River in part shade. The riverine habitat is comprised of a wide array of deciduous and evergreen trees, shrubs, and perennial plant species. Most of the plants along the river are adapted to acidic conditions and, while the aquatic habitat is generally fresh water, there is occasionally tidal or hurricane-caused saltwater intrusions. Slope: 0 percent. Aspect: North. The underlying geology of this area is characterized as Alluvium from undifferentiated Quaternary Sediments. Much of Florida's surface is covered by a varying thickness of undifferentiated sediments consisting of siliciclastics, organics and freshwater carbonates. Where these sediments exceed 20 feet (6.1 meters) thick, they were mapped as discrete units. To subdivide the undifferentiated sediments, those sediments occurring in flood plains were mapped as alluvial and flood plain deposits (Qal). Sediments showing surficial expression of beach ridges and dunes were mapped separately (Qbd) as were the sediments composing Trail Ridge (Qtr). Terrace sands were not mapped (refer to Healy [1975] for a discussion of the terraces in Florida). The subdivisions of the Undifferentiated Quaternary Sediments (Qu) are not lithostratigraphic units but are utilized to facilitate a better understanding of the State's geology. The siliciclastics are light gray, tan, brown to black, unconsolidated to poorly consolidated, clean to clayey, silty, unfossiliferous, variably organic-bearing sands to blue green to olive green, poorly to moderately consolidated, sandy, silty clays. Gravel is occasionally present in the panhandle. Organics occur as plant debris, roots, disseminated organic matrix and beds of peat. Freshwater carbonates, often referred to as marls in the literature, are scattered over much of the State. In southern Florida, freshwater carbonates are nearly ubiquitous in the Everglades. These sediments are buff colored to tan, unconsolidated to poorly consolidated, fossiliferous carbonate muds. Sand, silt and clay may be present in limited quantities. These carbonates often contain organics. The dominant fossils in the freshwater carbonates are mollusks. (Source: USGS Florida Geologic Map Data). The soils of this are characterized as Bibb-Kinston Series association. The Bibb Series consists of very deep, poorly drained, moderately permeable soils on flood plains of streams in the Southern Coastal Plain (133A) Major Land Resource Area. They formed in stratified loamy and sandy alluvium that are commonly and frequently flooded and water runs off the surface very slowly. Slopes range from 0 to 2 percent. Reaction ranges from extremely acid to strongly acid throughout. Content of mica flakes ranges from none to common. Content of rounded gravel typically ranges from 0 to 10 percent throughout, but may range to 50 percent in thin strata below a depth of 1 meter. Buried soil horizons, present in many pedons have the same range in color and texture as the surface horizon which has iron and organic matter concentrations in shades of brown and yellow ranging from none to common. Texture is sand, loamy sand, loamy fine sand, fine sandy loam, sandy loam, loam or silt loam. The Kinston Series consists of very deep, poorly drained, moderately permeable soils found on flood plains. They were formed in alluvial depositions. Slopes range from 0 to 2 percent. Soil reaction ranges from strongly acid to very strongly acid. Typical texture across the pedons is loamy, gravelly loam, sandy loam, and clay loam with dark concretions often present. (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions).Wild material2169929NA 88592
416NA 885932024RJLNJ-001Ilex glabra (L.) A. Gray New Jersey, United StatesNA2024COLLECTED11/24/2024Mullica Township: Both sides of Duerer Street, approximately 0.37 kilometers southeast of 4th Avenue.39.55885000, -74.6644250021Shade to almost full sun on compacted seasonally moist to temporarily wet areas of a mixed evergreen and deciduous woodland dominated by Pinus rigida. Vegetation of this area is very much defined by small changes in elevation that go from wet to very dry. Ilex glabra remains abundant throughout, though. Slope: N/A. Aspect: N/A. The underlying geology of this area is the Cohansey Formation of middle Miocene origin and comprised of sand, white to yellow with local gravel and clay. It is sometimes locally stained red or orange brown by iron oxides and (or) cemented into large blocks of ironstone. Unweathered clay is typically dark gray, but commonly weathers white where it is interbedded with thin beds of ironstone. Sand is typically medium grained and moderately sorted although it ranges from fine to very coarse grained and from poorly to well sorted. Sand consists of quartz and siliceous rock fragments. Some beds are locally micaceous. Maximum thickness in the map area is about 60 meters; however, thickness is difficult to determine because of the irregular basal contact and extensive post-depositional erosion. (Source: USGS New Jersey Geologic Map Data). The primary soil type in the area of this collection is classified as Mullica Series sandy loam, 0 to 2 percent slopes. These soils are typically coarse-loamy, siliceous, semiactive, and mesic typic humaquepts that are strongly to extremely acidic throughout the profile. The typical solum thickness ranges from 50-150+ cm. These soils are generally considered to be poorly drained. (Source: NRCS Web Soil Survey).Wild material2169930NA 88593
417NA 88479TXWL2024-029Ilex longipes Chapm. ex Trel. Texas, United StatesNA2024COLLECTED11/20/2024Little Cow Creek.31.06221000, -93.79603000Upland position above perennial creek.Wild material2169906NA 88479
418NA 88480TXWL2024-030Ilex coriacea (Pursh) Chapm. Texas, United StatesNANot Available2024COLLECTED11/20/2024Little Cow Creek.31.06280000, -93.79610000Upland position above perennial creek.Wild material2169907NA 88480
419NA 87836TRX241117-01VAIlex vomitoria Aiton Virginia, United StatesNA2024COLLECTED11/17/2024Eastern Shore of Virginia National Wildlife Refuge.37.13092000, -75.962530006Wild material2169889NA 87836
420NA 87838TRX241117-03NCIlex coriacea (Pursh) Chapm. North Carolina, United StatesNA2024COLLECTED11/17/2024Boiling Spring Lakes Community Center nature trail.34.02040000, -78.0646900016Wild material2169891NA 87838
421NA 87839TRX241117-04NCIlex cassine L. North Carolina, United StatesNA2024COLLECTED11/17/2024Boiling Spring Lakes Community Center nature trail.34.02040000, -78.0646900016Wild material2169892NA 87839
422NA 87840TRX241117-05NCIlex cassine L. North Carolina, United StatesNA2024COLLECTED11/17/2024Boiling Spring Lakes Community Center nature trail.34.02040000, -78.0646900016Wild material2169893NA 87840
423NA 87842TRX241117-08SCIlex opaca Aiton South Carolina, United StatesNA2024COLLECTED11/17/2024Francis Marion National Forest.33.15167000, -79.6025300031Wild material2169895NA 87842
424NA 87844TRX241118-02GAIlex cassine L. Georgia, United StatesNA2024COLLECTED11/18/2024St. Simons Island: Frederica Park.31.23766000, -81.356380006Wild material2169897NA 87844
425NA 87845TRX241119-01GAIlex coriacea (Pursh) Chapm. Georgia, United StatesNA2024COLLECTED11/19/2024St. Simons Island: Guale Preserve.31.21861000, -81.368720006Wild material2169898NA 87845
426NA 87848TRX241119-04GAIlex vomitoria Aiton Georgia, United StatesNA2024COLLECTED11/19/2024Jekyll Island.31.11344000, -81.406730004The first vegetation occuring at the top of the beach, where high tide is actively eroding.Wild material2169901NA 87848
427NA 88476TXWL2024-026Ilex verticillata (L.) A. Gray Texas, United StatesNANot Available2024COLLECTED11/12/2024Sabine River at I-10 at the Texas/Louisiana state line; north side of bridge.30.12830000, -93.70561000Bottomland.Wild material2169861NA 88476
428NA 88477TXWL2024-027Ilex ambigua Torr. Texas, United StatesNA2024COLLECTED11/12/2024Sabine River at I-10 at the Texas/Louisiana state line; south side of bridge.30.12654000, -93.70258000Bottomland.Wild material2169862NA 88477
429NA 88478TXWL2024-028Ilex vomitoria Aiton Texas, United StatesNA2024COLLECTED11/12/2024Big Thicket National Preserve: Turkey Creek Unit.: Pitcher Plant Trail.30.58602000, -94.34154000Bog/wetland.Wild material2169863NA 88478
430NA 87820'Blue Wave'Ilex hybr.Michigan, United StatesNANot Available2024DONATEDPRE 2004CultivarHolly Society of America test holly for 2024. Originated from the holly breeding program of the late Robert L. Tomayer (Wavecrest Nursery, Fennville, Michigan) and registered by his daughter, Carol T. Hop. The parent plant of this material was obtained by J. Resch directly from Mr. Tomayer at Wavecrest Nursery over 20 years ago.2169671NA 87820
431NA 88474JH2024NHIlex verticillata (L.) A. Gray New Hampshire, United StatesNANot Available2024COLLECTED10/18/2024North Conway, NH: near Saco River, river trail near Eastern Slope Camping Area [inferred; originally written, "River Trail Nr. East Slope."]Wild material2169440NA 88474
432NA 88475TXWL2024-025Ilex coriacea (Pursh) Chapm. Texas, United StatesNA2024COLLECTED10/16/2024Sabine National Forest C-142.Baygall below longleaf [pine] forest.Wild material2169441NA 88475
433NA 890802024RJLPA-006Ilex montana Torr. & A. Gray Pennsylvania, United StatesNA2024COLLECTED09/20/2024Berks and Schuylkill Counties: Hawk Mountain Sanctuary.40.64162200, -75.99122600418Plants are growing from shade to full sun on a dry-mesic to very dry, well-drained upland/montane habitat characterized by an upland mixed deciduous and evergreen forest that is very rocky. All the plants found in this habitat go through periods of drought during the growing season due to the limited litter/soil layer in which moisture can be held. Elevation: 405 to 430 meters. Slope: 5 to 25 percent. Aspect: various. Geology: The underlying geology of this area is of Silurian origin and comprised of Shawangunk Formation. This formation is primarily light- to dark-gray, fine- to very coarse-grained sandstone and conglomerate containing thin shale interbeds. (Source: USGS Pennsylvania Geologic Map Data). Soils: The surface soil profile of this area is characterized as the Hazleton Series which consists of deep and very deep, well drained soils formed in residuum of acid gray, brown or red sandstone on uplands. The matrix includes a considerable accumulation of thin, flat, coarse fragments of sandstone, limestone, or schist with diameters up to 15 cm. Slope ranges from 0-80%. Permeability is moderately rapid to rapid. (Source: USDA-NRCS Official Soil Series Descriptions). EPA Ecoregion (Level III): Ridge and Valley (67). EPA Ecoregion (Level IV): Northern Sandstone Ridges (67c).Wild material2169044NA 89080
434NA 890862024RJLPA-015Ilex verticillata (L.) A. Gray Pennsylvania, United StatesNA2024COLLECTED09/22/2024Outcrops along the Susquehanna River, accessed from the Historic Lock 12 parking lot off of Pennsylvania Road 372.39.81656600, -76.3313660038Plants are growing from part-sun to full sun in crevices of the outcrops and where organic debris has accumulated in between boulders along the Susquehanna River. The sites range from nearly xeric to dry mesic, at least on the surface. Plants are generally growing 4-10 m above the normal flow of the river; however, the Holtwood Dam is less than 1 kilometer upstream and debris found trapped in the boulders indicates that there is periodic flooding, at least, in the lower zone of 4+ meters or so above normal river flow. This habitat has a diverse assemblage of deciduous trees and shrubs as well as abundant herbaceous taxa. Slope: 0 to 10 percent. Aspect: Northeast (generally). Geology: The underlying geology of this area is probably of lower Paleozoic age and characterized as the Octoraro Formation. It includes albite-chlorite schist, phyllite, some hornblende gneiss, and granitized members. Soils: The area where this species occurs is right on the edge of two soil zones that transition from the river’s edge to mesic conditions upslope. The first is generally referred to as Rock Outcrop. This terrestrial matrix in which plants grow is largely exposed underlying rock geology with accumulated outwash, rock decomposition, and organic debris buildup. The second is Mt. Airy (80%) and Manor (20%) Series soils, 25 to 60 percent slopes, extremely stony (MRF). (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions). EPA Ecoregion (Level III): Northern Piedmont (64). EPA Ecoregion (Level IV): Piedmont Uplands (64c).Wild material2169050NA 89086
435NA 890902024RM-TK-003Ilex longipes Chapm. ex Trel. Alabama, United StatesNA2024COLLECTED09/19/2024Woodland slopes around and below the hilltop parking area leading down to the Silver Creek Lake Campground off of Silver Creek Lake Road.31.66612294, -87.5756986039Plants are growing in part-shade to shade of a moist mesic deciduous and evergreen woodland on dry, well-drained soil to an area 1-2 meters above a small seasonally spring-fed creek at 31.664428°, -87.574614°. Slope: 5 to 10 percent. Aspect: Southeast. Geology: The underlying geology of this area is Claiborne Group, Tallahatta Formation, of Eocene origin, with white to very light-greenish-gray thin-bedded to massive siliceous claystone; interbedded with thin layers of fossiliferous clay, sandy clay, and glauconitic sand and sandstone. (Source: USGS Alabama Geologic Map Data). Soils: The primary soil types in the collection area are classified as Arundel-Cantuche Complex soils (15 to 35 percent slopes). The Arundel Series consists of moderately deep, well drained, very slowly permeable soils on dissected uplands of the Southern Coastal Plain. They formed in marine deposits consisting of acid clays underlain by horizontally bedded sandstone, buhrstone and siltstone. These soils are strongly acid. Slopes range from 2 to 60 percent. The Cantuche Series consists of shallow, well drained, moderately permeable soils that formed over horizontally bedded claystone. These soils are on narrow ridgetops and narrow benches on steep side-slopes of dissected landscapes in the Coastal Plain. They are strongly acid. Slopes range from 2 to 35 percent. (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions). EPA Ecoregion (Level III): Southeastern Plains (65). EPA Ecoregion (Level IV): Buhrstone/Lime Hills (65q).Wild material2169054NA 89090
436NA 890922024RM-TK-005Ilex amelanchier M. A. Curtis ex Chapm. Florida, United StatesNA2024COLLECTED09/29/2024Yellow River Water Management Area: found in an area where an unnamed, unpaved road dead ends near the Yellow River. This unpaved road is accessed from Old River Road to the east near the junction with Shockley Springs Road. Plants are accessed by foot along the edge of the river and associated wetland depressions to the west of the road.30.86946200, -86.5934900023Plants growing in seasonally inundated swamp habitats associated with the Yellow River. Plants are emergent during portions of the year, primarily during the growing season. During this time of collection, water had receded from the wetlands but the area was still quite wet with many tiny stream rivulets still draining. Slope: 0 percent. Aspect: N/A. Geology: The underlying geology of this area is the Alum Bluff Group of Miocene origin. It includes the Chipola Formation, Oak Grove Sand, Shoal River Formation, Choctawhatchee Formation and the Jackson Bluff Formation. The formations included in this group are generally defined based on their molluscan faunas and stratigraphic position. Puri (1953) described sediment facies as they relate to the formations of the Alum Bluff Group. These sediments are lithologically distinct as a group, not as individual units. The Alum Bluff Group crops out or is beneath a thin overburden in the western panhandle from river valleys in Okaloosa County eastward to western Jackson County. The Alum Bluff Group consists of clays, sands and shell beds which may vary from fossiliferous, sandy clays to unfossiliferous sands and clays and occasional carbonate beds. Mica is a common constituent and glauconite and phosphate occur sporadically. Induration varies from essentially nonindurated in sands to well indurated in carbonate lenses. Colors range from cream to olive gray with mottled reddish brown in weathered sections. Sand grain size varies from very fine to very coarse with sporadic occurrences of gravel. These sediments generally have low permeabilities and are part of the intermediate confining unit/aquifer system. (Source: USGS Florida Geologic Map Data). Soils: The surface soil profile of this area is primarily Kinston, Johnston, and Bibb soils, frequently flooded. Kinston Series soils are poorly drained, alluvial, fine-loamy, siliceous, and acid soils of flood plains. They are peaty to loamy friable, slightly sticky and slightly plastic in the upper 0 to 7.5 cm of the strata with less sand and more silt-loam in progressively deeper strata. The Johnston Series consists of very deep, very poorly drained coarse-loamy, siliceous, active, acid, thermic Cumulic Humaquepts. The upper 0-75 cm is black, mucky loam that is friable and very strongly acid. The lower strata to 1.75 m are dark gray, loamy fine sand, single grained, loose and very strongly a).id. The Bibb Series soils consist of very deep, poorly drained, moderately permeable soils on flood plains of streams in the Southern Coastal Plain (133A) Major Land Resource Area. They formed in stratified loamy and sandy alluvium that are commonly and frequently flooded and water runs off the surface very slowly. Slopes range from 0 to 2 percent. (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions). EPA Ecoregion (Level III): Southeastern Plains (65). EPA Ecoregion (Level IV): Southeastern Floodplains and Low Terraces (65p).Wild material2169056NA 89092
437NA 890932024RM-TK-006Ilex ambigua Torr. Florida, United StatesNA2024COLLECTED09/29/2024Blackwater River Wildlife Management Area: growing on the south side of Bud Bass Road, approximately 0.3 kilometers east of Training Center R./Norma Riley Rd.30.75759720, -86.8018722029This plant is growing in part-shade under mesic conditions along the edge of a longleaf pine plantation. Slope: 0 percent. Aspect: North. Geology: The underlying geology of this area is probably of lower Pliocene age and characterized as the Citronelle Formation. The Citronelle Formation is widespread in the Gulf Coastal Plain. The type section for the Citronelle Formation, named by Matson (1916), is near Citronelle, Alabama. The Citronelle Formation grades laterally, through a broad facies transition, into the Miccosukee Formation of the eastern Florida panhandle. Coe (1979) investigated the Citronelle Formation in portions of the western Florida panhandle. The Citronelle Formation is a siliciclastic, deltaic deposit that is lithologically similar to, and time equivalent with, the Cypresshead Formation and, at least in part, the Long Key Formation (Cunningham et al., 1998) of the peninsula. In the western panhandle, some of the sediments mapped as Citronelle Formation may be reworked Citronelle. The lithologies are the same and there are few fossils present to document a possible younger age. The Citronelle Formation consists of gray to orange, often mottled, unconsolidated to poorly consolidated, very fine to very coarse, poorly sorted, clean to clayey sands. It contains significant amounts of clay, silt and gravel which may occur as beds and lenses and may vary considerably over short distances. Limonite nodules and limonite-cemented beds are common. Marine fossils are rare but fossil pollen, plant remains and occasional vertebrates are found. Much of the Citronelle Formation is highly permeable. It forms the Sand and Gravel Aquifer of the surficial aquifer system. (Source: USGS Florida Geologic Map Data). Soils: The soil of this area is characterized as Lakeland Series Sand, 0 to 5 percent slopes (21). The Lakeland Series consists of very deep, excessively drained, rapid to very rapidly permeable soils on uplands. They formed in thick beds of eolian or marine and/or fluvio-marine sands in the Southern Coastal Plain MLRA (133A), the Carolina and Georgia Sandhills (MLRA 137), the Eastern Gulf Coast Flatwoods (MLRA 152A) and the Atlantic Coast Flatwoods (MLRA 153A). (Source: California Soil Resource Lab/NRCS Official Soil Series Descriptions). EPA Ecoregion (Level III): Southeastern Plains (65). EPA Ecoregion (Level IV): Southern Pine Plains and Hills (65f).Wild material2169057NA 89093
438NA 87823FL24-01Ilex cassine L. Florida, United StatesNA2024COLLECTED09/05/2024Matanzas State Forest.29.72143991, -81.28386478Boggy blackwater ditches along edge of dirt road. EPA Ecoregion III: Southern Coastal Plain (75). EPA Ecoregion IV: Eastern Florida Flatwoods (75d). Geology: Beach Ridge and Dune. Soil: Immokalee Series Fine Sand.Wild material2168973NA 87823
439NA 87824FL24-02Ilex cassine L. Florida, United StatesNA2024COLLECTED09/05/2024Wickham Park.28.17068273, -80.66140942A very open, sandy, and xeric forest that rapidly turns wet and boggy where P. palustris (and Ilex cassine) occurs. EPA Ecoregion III: Southern Coastal Plain (75). EPA Ecoregion IV: Eastern Florida Flatwoods (75d). Geology: Anastasia Formation. Soil: Myakka Series Sand, 0 to 2 percent slopes.Wild material2168974NA 87824
440NA 87825FL24-03Ilex ambigua Torr. Florida, United StatesNA2024COLLECTED09/07/2024Rye Preserve, along the Manatee River.27.50951673, -82.35916675Open scrub, xeric, sandy. EPA Ecoregion III: Southern Coastal Plain (75). EPA Ecoregion IV: Southwestern Florida Flatwood (75b). Geology: Hawthorn Group, Peace River Formation. Soil: Cassia Series Fine Sand, moderately well drained.Wild material2168975NA 87825
441NA 87828GA24-01Ilex glabra (L.) A. Gray Georgia, United StatesNANot Available2024COLLECTED09/09/2024Crooked River State Park.30.83654580, -81.55628447Longleaf Pine flatwoods, very wet. EPA Ecoregion III: Southern Coastal Plain (75). Geology: Princess Anne Shoreline Complex-Barrier Island Facies. EPA Ecoregion IV: Sea Islands/Coastal Marsh (75j). Soil: Mandarin Series Fine Sand, 0 to 2 percent slopes.Wild material2168978NA 87828
442NA 87834TRX240901-02WVIlex montana Torr. & A. Gray West Virginia, United StatesNA2024COLLECTED09/01/2024Blackwater Falls State Park.39.10665000, -79.48680000967Wild material2168853NA 87834
443NA 87687'Lacquerberry'Ilex opaca Aiton NANot Available2024CultivarOne own root, potted plant received.2167509NA 87687
444NA 87721'Jim Dandy'Ilex verticillata (L.) A. Gray NANot Available2024CultivarFull name as recorded at the U.S. National Arboretum: Ilex verticillata 'Jim Dandy'.2192965NA 87721
445NA 88852TJK2023FL-047Ilex glabra (L.) A. Gray Florida, United StatesNA2024COLLECTED12/26/2023South of Highway 98, west of Twine Road. Track has a very large powerline right-of-way to the east, and George Washington Swamp bottom to the west.29.77640000, -85.274100006Longleaf Pine-Saw Palmetto. This area appears to be declining in quality, as witnessed by the declining in Rhododendron stem/plant density. Possible contributing issues: lacking necessary fire regime, has not seen fire in 8-10 years, encroaching woody shrub layer, accumulating organic matter (pine needle), expanding tree canopy, Hurricane Michael. Slope: less than 7%. Aspect: West. Elevation: 20 feet.Wild material2165707NA 88852
446NA 88827RJL-RM2023FLAL-001Ilex ambigua Torr. Florida, United StatesNA2023COLLECTED12/05/2023Yellow River Wildlife Management Area, along DOF [Department of Forestry] Y9 dirt road 0.08 kilometers east of Miller Bluff Road.30.65289100, -86.8760280044Growing in part shade to sun in very dry, highly competitive site still recovering from the logging of pines in the past decade. This young woodland is comprised of a combination of evergreen and deciduous species competing for available resources and light before the young sand pines begin to dominate the upper canopy. Slope: 0 percent. Aspect: N/A. The underlying geology of this area is the Citronelle Formation, of Pliocene origin, and consists of gray to orange, often mottled, unconsolidated to poorly consolidated, very fine to very coarse, poorly sorted, clean to clayey sands. It contains significant amounts of clay, silt and gravel which may occur as beds and lenses and may vary considerably over short distances. Limonite nodules and limonite-cemented beds are common. (Source: USGS Florida Geologic Map Data.) The primary soil type in the collection area is classified as Lakeland Sand soils (0 to 5 percent slopes). The surface soil profile of this area is primarily the Lakeland series which consists of very deep, excessively drained, rapid to very rapidly permeable soils on uplands. They formed in thick beds of eolian or marine and/or fluvio-marine sands. Thickness of the sand exceeds 2 meters. Silt plus clay in the upper meter ranges from 5-10 percent. Reaction ranges from very strongly acid to moderately acid throughout. (Source: NRCS Web Soil Survey.)Wild material2165657NA 88827
447NA 88833RJL-RM2023FLAL-007Ilex decidua Walter Florida, United StatesNANot Available2023COLLECTED12/06/2023Plants are growing on the north side of Oil Plant Road between the road and Bray Mill Creek (which dumps into the nearby Escambia River), 0.37 kilometers east of McDavid Park.30.97645800, -87.1856260037Growing in full sun to part shade in seasonally moist to wet areas associated with Bray Mill Creek; however, during low rainfall periods, plants are in dry soil. The habitat is heavily degraded due to logging of most of the area except a narrow woodland along the creek edge. The gravel road also causes significant run off and deposition of road debris. Despite this fact, the Ilex decidua is thriving. Slope: 0 to 5 percent. Aspect: North to northeast. The underlying geology of this area is classified as Alluvium, of Pleistocene/Holocene origin and consists of undifferentiated Quaternary sediments of varying thickness including siliciclastics, organics and freshwater carbonates. The siliciclastics are light gray, tan, brown to black, unconsolidated to poorly consolidated, clean to clayey, silty, unfossiliferous, variably organic-bearing sands to blue green to olive green, poorly to moderately consolidated, sandy, silty clays. Organics occur as plant debris, roots, disseminated organic matrix and beds of peat. Freshwater carbonates, often referred to as marls in the literature, are scattered. (Source: USGS Florida Geologic Map Data.) The surface soil profile of this area is primarily Chewacla-Wahee-Riverview Series Association. The Chewacla Series consists of very deep, somewhat poorly drained, fine loamy alluvial soils of flood plains that range from strongly to very strongly acid. These soils contain very little rock and can reach a thickness of 2 m. Wahee Series consists of very deep, somewhat poorly drained, clayey and loamy marine or fluviomarine sediments. These soils are strongly to very strongly acid. The Riverview Series consists of very deep, well drained, moderately permeable soils that formed in loamy alluvium on flood plains. These soils contain a slightly higher level of mica and iron deposits and range from slightly to strongly acid. (Source: NRCS Web Soil Survey.)Wild material2165663NA 88833
448NA 88838RJL-RM2023FLAL-012Ilex opaca Aiton Florida, United StatesNA2023COLLECTED12/08/2023North side of Sid Hayes Road approximately 65 m southeast of the junction with Watson Road.30.83277400, -87.2432450032Growing in shade, part shade and light shade in dry mesic to moist, occasionally wet, habitats along the edge of the road and in the woodland interior. This species is remarkably adaptable. The only drawback to the range of habitats is that in deep shade, very few fruit can be found, presumably because there is far less flower production and/or fewer pollinators. Slope: 0 to 3 percent. Aspect: South. The underlying geology of this area is the Citronelle Formation of Pliocene origin and consists of gray to orange, often mottled, unconsolidated to poorly consolidated, very fine to very coarse, poorly sorted, clean to clayey sands. It contains significant amounts of clay, silt and gravel which may occur as beds and lenses and may vary considerably over short distances. Limonite nodules and limonite-cemented beds are common. (Source: USGS Florida Geologic Map Data.) The surface soil profile of this area is Dothan Series fine sandy loam soils (5 to 8 percent slopes). The Dothan Series consists of very deep, well drained soils that formed in thick beds of unconsolidated, medium to fine-textured marine sediments. Dothan soils are on interfluves. The surface 0-33 cm is characterized by sandy-loam, well-drained soils that are strongly acid. From 33-200 cm they are characterized by sandy clay loam soils that are more moisture retentive and strongly acid. (Source: NRCS Web Soil Survey.)Wild material2165668NA 88838
449NA 88845RJL-RM2023FLAL-019Ilex opaca Aiton Florida, United StatesNA2023COLLECTED12/13/2023Yellow River Water Management Area, found along an unnamed unpaved road leading down to the Yellow River. This unpaved road is accessed from Old River Road to the east near the junction with Shockley Springs Road.30.87382600, -86.6022130035Plants are growing along the road on the woodland edge on a small bluff above a creek in well-drained to dry habitat in part shade. They are also found sporadically in areas along the Yellow River where occasional inundation occurs. Slope: N/A. Aspect: N/A. The underlying geology of this area is the Alum Bluff Group of Miocene origin. It includes the Chipola Formation, Oak Grove Sand, Shoal River Formation, Choctawhatchee Formation and the Jackson Bluff Formation. The formations included in this group are generally defined on the basis of their molluscan faunas and stratigraphic position. Puri (1953) described sediment facies as they relate to the formations of the Alum Bluff Group. These sediments are lithologically distinct as a group, not as individual units. The Alum Bluff Group crops out or is beneath a thin overburden in the western panhandle from river valleys in Okaloosa County eastward to western Jackson County. The Alum Bluff Group consists of clays, sands and shell beds which may vary from fossiliferous, sandy clays to unfossiliferous sands and clays and occasional carbonate beds. Mica is a common constituent and glauconite and phosphate occur sporadically. Induration varies from essentially nonindurated in sands to well indurated in carbonate lenses. Colors range from cream to olive gray with mottled reddish brown in weathered sections. Sand grain size varies from very fine to very coarse with sporadic occurrences of gravel. These sediments generally have low permeabilities and are part of the intermediate confining unit/aquifer system. (Source: USGS Florida Geologic Map Data.) The surface soil profile of this area is primarily Bonifay-Dothan-Angie complex (5 to 12 percent slopes). Bonifay soil consists of very deep, well drained, moderately slowly permeable soils on ridges and side slopes in the Southern Coastal Plain. They formed in thick beds of sandy and loamy marine sediments. Solum thickness ranges from 1.5-2.0 m. Soil reaction ranges from very strongly acid to moderately acid throughout, except where the surface has been limed. Ironstone pebbles, 2 to 15 mm in size, range from 0 to 5 percent, by volume. Depth to the plinthic horizon ranges from 1.1-1.7 m. Some horizons contain up to 25 percent plinthite by volume. The Dothan Series consists of very deep, well drained soils that formed in thick beds of unconsolidated, medium to fine-textured marine sediments. Dothan soils are on interfluves. The surface 0-33 cm is characterized by sandy-loam, well-drained soils that are strongly acid. From 33-200 cm they are characterized by sandy clay loam soils that are more moisture retentive and strongly acid. The Angie series consists of very deep, moderately well drained, slowly permeable soils that formed in loamy and clayey sediment on Tertiary-aged uplands. These soils are on broad, level to strongly sloping coastal plains. Slope is dominantly 1 to 5 percent, but the range is 0 to 12 percent. (Source: NRCS Web Soil Survey.)Wild material2165675NA 88845
450NA 88846RJL-RM2023FLAL-020Ilex decidua Walter Florida, United StatesNA2023COLLECTED12/13/2023Yellow River Water Management Area: found in an area where an unnamed unpaved road dead ends near the Yellow River. This unpaved road is accessed from Old River Road to the east near the junction with Shockley Springs Road.30.86973100, -86.5916870023Plants are growing primarily in part-shade to part-sun in low-lying moist habitats associated with the Yellow River. They are in the flood zone, but not regularly inundated. Slope: N/A. Aspect: N/A. The underlying geology of this area is the Alum Bluff Group of Miocene origin. It includes the Chipola Formation, Oak Grove Sand, Shoal River Formation, Choctawhatchee Formation and the Jackson Bluff Formation. The formations included in this group are generally defined on the basis of their molluscan faunas and stratigraphic position. Puri (1953) described sediment facies as they relate to the formations of the Alum Bluff Group. These sediments are lithologically distinct as a group, not as individual units. The Alum Bluff Group crops out or is beneath a thin overburden in the western panhandle from river valleys in Okaloosa County eastward to western Jackson County. The Alum Bluff Group consists of clays, sands and shell beds which may vary from fossiliferous, sandy clays to unfossiliferous sands and clays and occasional carbonate beds. Mica is a common constituent and glauconite and phosphate occur sporadically. Induration varies from essentially nonindurated in sands to well indurated in carbonate lenses. Colors range from cream to olive gray with mottled reddish brown in weathered sections. Sand grain size varies from very fine to very coarse with sporadic occurrences of gravel. These sediments generally have low permeabilities and are part of the intermediate confining unit/aquifer system. (Source: USGS Florida Geologic Map Data.) The surface soil profile of this area is primarily Kinston, Johnston, and Bibb soils, frequently flooded. Kinston Series soils are poorly drained, alluvial, fine-loamy, siliceous, and acid soils of flood plains. They are peaty to loamy friable, slightly sticky and slightly plastic in the upper 0 to 7.5 cm of the strata with less sand and more silt-loam in progressively deeper strata. The Johnston Series consists of very deep, very poorly drained coarse-loamy, siliceous, active, acid, thermic Cumulic Humaquepts. The upper 0-75 cm is black, mucky loam that is friable and very strongly acid. The lower strata to 1.75 m is dark gray, loamy fine sand, single grained, loose and very strongly acid. The Bibb Series soils consists of very deep, poorly drained, moderately permeable soils on flood plains of streams in the Southern Coastal Plain (133A) Major Land Resource Area. They formed in stratified loamy and sandy alluvium that are commonly and frequently flooded and water runs off the surface very slowly. Slopes range from 0 to 2 percent. (Source: NRCS Web Soil Survey.)Wild material2165676NA 88846
451NA 88847RJL-RM2023FLAL-021Ilex amelanchier M. A. Curtis ex Chapm. Florida, United StatesNA2023COLLECTED12/13/2023Yellow River Water Management Area, found in an area where an unnamed unpaved road dead ends near the Yellow River. This unpaved road is accessed from Old River Road to the east near the junction with Shockley Springs Road. Individuals are accessed by foot along the edge of the river and associated wetland depressions to the west of the road.30.86980100, -86.5921050023Plants growing in seasonally inundated swamp habitats associated with the Yellow River. Plants are emergent during portions of the year, primarily during the growing season. During this time of collection, water had receded from the wetlands but the area was still quite wet with many tiny stream rivulets still draining. Slope: N/A. Aspect: N/A. The underlying geology of this area is the Alum Bluff Group of Miocene origin. It includes the Chipola Formation, Oak Grove Sand, Shoal River Formation, Choctawhatchee Formation and the Jackson Bluff Formation. The formations included in this group are generally defined on the basis of their molluscan faunas and stratigraphic position. Puri (1953) described sediment facies as they relate to the formations of the Alum Bluff Group. These sediments are lithologically distinct as a group, not as individual units. The Alum Bluff Group crops out or is beneath a thin overburden in the western panhandle from river valleys in Okaloosa County eastward to western Jackson County. The Alum Bluff Group consists of clays, sands and shell beds which may vary from fossiliferous, sandy clays to unfossiliferous sands and clays and occasional carbonate beds. Mica is a common constituent and glauconite and phosphate occur sporadically. Induration varies from essentially nonindurated in sands to well indurated in carbonate lenses. Colors range from cream to olive gray with mottled reddish brown in weathered sections. Sand grain size varies from very fine to very coarse with sporadic occurrences of gravel. These sediments generally have low permeabilities and are part of the intermediate confining unit/aquifer system. (Source: USGS Florida Geologic Map Data.) The surface soil profile of this area is primarily Kinston, Johnston, and Bibb soils, frequently flooded. Kinston Series soils are poorly drained, alluvial, fine-loamy, siliceous, and acid soils of flood plains. They are peaty to loamy friable, slightly sticky and slightly plastic in the upper 0 to 7.5 cm of the strata with less sand and more silt-loam in progressively deeper strata. The Johnston Series consists of very deep, very poorly drained coarse-loamy, siliceous, active, acid, thermic Cumulic Humaquepts. The upper 0-75 cm is black, mucky loam that is friable and very strongly acid. The lower strata to 1.75 m is dark gray, loamy fine sand, single grained, loose and very strongly acid. The Bibb Series soils consists of very deep, poorly drained, moderately permeable soils on flood plains of streams in the Southern Coastal Plain (133A) Major Land Resource Area. They formed in stratified loamy and sandy alluvium that are commonly and frequently flooded and water runs off the surface very slowly. Slopes range from 0 to 2 percent. (Source: NRCS Web Soil Survey.)Wild material2165677NA 88847
452NA 88848RJL-RM2023FLAL-022Ilex amelanchier M. A. Curtis ex Chapm. Florida, United StatesNANot Available2023COLLECTED12/13/2023Yellow River Water Management Area, found in an area where an unnamed unpaved road dead ends near the Yellow River. This unpaved road is accessed from Old River Road to the east near the junction with Shockley Springs Road. Individuals are accessed by foot along the edge of the river and associated wetland depressions to the west of the road.30.86945000, -86.5934880023Plants growing in seasonally inundated swamp habitats associated with the Yellow River. Plants are emergent during portions of the year, primarily during the growing season. During this time of collection, water had receded from the wetlands but the area was still quite wet with many tiny stream rivulets still draining. Slope: N/A. Aspect: N/A. The underlying geology of this area is the Alum Bluff Group of Miocene origin. It includes the Chipola Formation, Oak Grove Sand, Shoal River Formation, Choctawhatchee Formation and the Jackson Bluff Formation. The formations included in this group are generally defined on the basis of their molluscan faunas and stratigraphic position. Puri (1953) described sediment facies as they relate to the formations of the Alum Bluff Group. These sediments are lithologically distinct as a group, not as individual units. The Alum Bluff Group crops out or is beneath a thin overburden in the western panhandle from river valleys in Okaloosa County eastward to western Jackson County. The Alum Bluff Group consists of clays, sands and shell beds which may vary from fossiliferous, sandy clays to unfossiliferous sands and clays and occasional carbonate beds. Mica is a common constituent and glauconite and phosphate occur sporadically. Induration varies from essentially nonindurated in sands to well indurated in carbonate lenses. Colors range from cream to olive gray with mottled reddish brown in weathered sections. Sand grain size varies from very fine to very coarse with sporadic occurrences of gravel. These sediments generally have low permeabilities and are part of the intermediate confining unit/aquifer system. (Source: USGS Florida Geologic Map Data.) The surface soil profile of this area is primarily Kinston, Johnston, and Bibb soils, frequently flooded. Kinston Series soils are poorly drained, alluvial, fine-loamy, siliceous, and acid soils of flood plains. They are peaty to loamy friable, slightly sticky and slightly plastic in the upper 0 to 7.5 cm of the strata with less sand and more silt-loam in progressively deeper strata. The Johnston Series consists of very deep, very poorly drained coarse-loamy, siliceous, active, acid, thermic Cumulic Humaquepts. The upper 0-75 cm is black, mucky loam that is friable and very strongly acid. The lower strata to 1.75 m is dark gray, loamy fine sand, single grained, loose and very strongly acid. The Bibb Series soils consists of very deep, poorly drained, moderately permeable soils on flood plains of streams in the Southern Coastal Plain (133A) Major Land Resource Area. They formed in stratified loamy and sandy alluvium that are commonly and frequently flooded and water runs off the surface very slowly. Slopes range from 0 to 2 percent. (Source: NRCS Web Soil Survey.)Wild material2165678NA 88848
453NA 88849RJL-RM2023FLAL-023Ilex amelanchier M. A. Curtis ex Chapm. Florida, United StatesNANot Available2023COLLECTED12/13/2023Yellow River Water Management Area, found in an area where an unnamed unpaved road dead ends near the Yellow River. This unpaved road is accessed from Old River Road to the east near the junction with Shockley Springs Road. Individuals are accessed by foot along the edge of the river and associated wetland depressions to the west of the road.30.86982300, -86.5937830023Plants growing in seasonally inundated swamp habitats associated with the Yellow River. Plants are emergent during portions of the year, primarily during the growing season. During this time of collection, water had receded from the wetlands but the area was still quite wet with many tiny stream rivulets still draining. Slope: N/A. Aspect: N/A. The underlying geology of this area is the Alum Bluff Group of Miocene origin. It includes the Chipola Formation, Oak Grove Sand, Shoal River Formation, Choctawhatchee Formation and the Jackson Bluff Formation. The formations included in this group are generally defined on the basis of their molluscan faunas and stratigraphic position. Puri (1953) described sediment facies as they relate to the formations of the Alum Bluff Group. These sediments are lithologically distinct as a group, not as individual units. The Alum Bluff Group crops out or is beneath a thin overburden in the western panhandle from river valleys in Okaloosa County eastward to western Jackson County. The Alum Bluff Group consists of clays, sands and shell beds which may vary from fossiliferous, sandy clays to unfossiliferous sands and clays and occasional carbonate beds. Mica is a common constituent and glauconite and phosphate occur sporadically. Induration varies from essentially nonindurated in sands to well indurated in carbonate lenses. Colors range from cream to olive gray with mottled reddish brown in weathered sections. Sand grain size varies from very fine to very coarse with sporadic occurrences of gravel. These sediments generally have low permeabilities and are part of the intermediate confining unit/aquifer system. (Source: USGS Florida Geologic Map Data.) The surface soil profile of this area is primarily Kinston, Johnston, and Bibb soils, frequently flooded. Kinston Series soils are poorly drained, alluvial, fine-loamy, siliceous, and acid soils of flood plains. They are peaty to loamy friable, slightly sticky and slightly plastic in the upper 0 to 7.5 cm of the strata with less sand and more silt-loam in progressively deeper strata. The Johnston Series consists of very deep, very poorly drained coarse-loamy, siliceous, active, acid, thermic Cumulic Humaquepts. The upper 0-75 cm is black, mucky loam that is friable and very strongly acid. The lower strata to 1.75 m is dark gray, loamy fine sand, single grained, loose and very strongly acid. The Bibb Series soils consists of very deep, poorly drained, moderately permeable soils on flood plains of streams in the Southern Coastal Plain (133A) Major Land Resource Area. They formed in stratified loamy and sandy alluvium that are commonly and frequently flooded and water runs off the surface very slowly. Slopes range from 0 to 2 percent. (Source: NRCS Web Soil Survey.)Wild material2165679NA 88849
454NA 88818TJK2023FL-037Ilex opaca Aiton var. arenicola (Ashe) Ashe Florida, United StatesNANot Available2023COLLECTED12/05/2023Lake Wales Ridge State Forest.27.68579700, -81.4142250027Xeric scrub. Slope: flat. Aspect: Southeast. Elevation: 89 feet.Wild material2165647NA 88818
455NA 88824TJK2023FL-041Ilex opaca Aiton Florida, United StatesNA2023COLLECTED12/11/2023Holmes Creek, Live Oak Landing.30.60450000, -85.766700006Bottomland floodpain of Holmes Creek. Slope: flat. Aspect: Southwest. Elevation: 20 feet.Wild material2165653NA 88824
456NA 88825TJK2023FL-042Ilex verticillata (L.) A. Gray Florida, United StatesNA2023COLLECTED12/11/2023Holmes Creek, Live Oak Landing.30.60450000, -85.766700006Bottomland floodpain of Holmes Creek. Slope: flat. Aspect: Southwest. Elevation: 20 feet.Wild material2165654NA 88825
457NA 88826TJK2023FL-043Ilex decidua Walter Florida, United StatesNA2023COLLECTED12/14/2023Upper Escambia River.30.87870000, -87.291200009Hammock, bench-like river floodplain/bottomland. Slope: flat. Aspect: East-southeast. Elevation: 30 feet.Wild material2165655NA 88826
458NA 88811TXWL2023-017Ilex longipes Chapm. ex Trel. Texas, United StatesNANot Available2023COLLECTED12/04/2023Indian Hills Community/Wiergate: County Road 1089 roadside.31.00317700, -93.72888600Roadside. Upland; sandy loam soil.Wild material2165614NA 88811
459NA 88812TXWL2023-018Ilex ambigua Torr. Texas, United StatesNA2023COLLECTED12/05/2023Davy Crockett National Forest: Forest Service Road 526 roadside.31.49017400, -95.22343300Roadside. Shortleaf mixed upland hardwood forest; dry-mesic.Wild material2165615NA 88812
460NA 88813TXWL2023-020Ilex vomitoria Aiton Texas, United StatesNA2023COLLECTED12/06/2023Private property (Bayou Bleu Farm).31.66376880, -94.01070880Road edge of wooded road; mixed pine-hardwood stand.Wild material2165617NA 88813
461NA 88814TXWL2023-021Ilex ambigua Torr. Texas, United StatesNA2023COLLECTED12/09/2023Roadside between Big Cow Creek and Hardy Cemetery.31.03548900, -93.85086800Roadside; mesic, mixed hardwood/pine forest.Wild material2165618NA 88814
462NA 88815TXWL2023-022Ilex opaca Aiton Texas, United StatesNA2023COLLECTED12/10/2023Angelina National Forest: Upland Island Wilderness Area: Neches River bottom.31.03919400, -94.37937900Bottomland hardwood forest.Wild material2165619NA 88815
463NA 88816TXWL2023-023Ilex decidua Walter Texas, United StatesNA2023COLLECTED12/10/2023Angelina National Forest: Upland Island Wilderness Area: Neches River bottom.31.03478500, -94.38286600Bottomland hardwood forest.Wild material2165620NA 88816
464NA 88802TJK2023LA/TX-002Ilex decidua Walter Louisiana, United StatesNA2023COLLECTED11/21/2023Interstate 20, westbound, approximately 6 miles west of U.S. Highway 65 and 7.5 miles east of Louisiana Highway 577. Tensas River watershed.32.41950000, -91.2967000021Damp wooded treeline. Slope: 12%. Aspect: West-southwest. Elevation: 69 feet.Wild material2159185NA 88802
465NA 88809TJK2023LA/TX-009Ilex decidua Walter Texas, United StatesNA2023COLLECTED11/27/2023Southbound Highway 281, west side of road, approximately 13 miles south of Highway 290 and 1 mile north of RR [rural route] 473 East.30.01610000, -98.40610000407Roadside, brushy fencerow. Slope: 10%. Aspect: East-northeast. Elevation: 1336 feet.Wild material2159192NA 88809
466NA 88800TXWL2023-015Ilex longipes Chapm. ex Trel. Texas, United StatesNA2023COLLECTED11/15/2023Sabine National Forest: Compartment 65 along County Road 131 near White Rock, Texas.31.55793500, -94.01980900Upland; mixed hardwood and pine; sandy soil; rocky outcrops.Wild material2159157NA 88800
467NA 88690RJL2023NJ-003Ilex verticillata (L.) A. Gray New Jersey, United StatesNA2023COLLECTED11/10/2023Franklin Township: Piney Hollow Preservation Area (the site is owned by Franklin Township). This species was found below the dam at the far end of McCarthys Lakes. Plants accessed by foot along trails throughout property.39.57407200, -74.9251860028Plants growing in part-shade to sun of the moist to saturated areas below the McCarthys Lakes dam in the wetlands associated with the New Squankum Branch. The area appears to be inundated year-round with plants growing on low mounds surrounded by obligate wetland species. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Middle Miocene, Serravallian origin and characterized as an Cohansey Formation. It is comprised of sand, white to yellow with local gravel and clay. Locally stained red or orange brown by iron oxides and (or) cemented into large blocks of ironstone. Unweathered clay is typically dark gray, but commonly weathers white where interbedded with thin beds of ironstone. Unit is a complex of interfingering marine and nonmarine facies. Sand is typically medium grained and moderately sorted although it ranges from fine to very coarse grained and from poorly to well sorted. Sand consists of quartz and siliceous rock fragments. Some beds are locally micaceous. In general, the sand is crossbedded, although the style of crossbedding varies significantly with the paleoenvironment. Trough crossbedding predominates, especially in the nonmarine channel fill deposits, and the scale of the crossbeds varies from small to large. Maximum thickness in the map area is about 60 m; however, thickness is difficult to determine because of the irregular basal contact and extensive post-depositional erosion. There is as much as 18 m of relief along the basal contact. The basal contact is sharp, undulatory, and directly overlain by a thin gravel bed. The Cohansey Formation unconformably overlies the Kirkwood Formation and is found in channels cut down into the Kirkwood. Where the Kirkwood consists of sandy, light-colored sediments, the basal contact of the Cohansey is drawn below crossbedded sediments. Where the Kirkwood consists of dark-colored silty beds, the basal contact is drawn between light-colored Cohansey sediments and the underlying dark-colored sediments. The Cohansey was markedly thinned because of erosion prior to deposition of overlying units in the western and southern parts of the southern sheet. The unit has been extensively eroded and stripped from large areas of the New Jersey Coastal Plain, particularly in the central sheet where outliers are common. In spite of its widespread nature, the Cohansey is poorly exposed because of its loose sandy composition, which causes it to erode easily. (Source: USGS New Jersey Geologic Map Data.) The primary soil type in the collection area is classified as Manahawkin Series muck, 0 to 2 percent slopes, frequently flooded, Northern Coastal Plain. Manahawkin Series soils are very deep, very poorly drained soils that are sandy or sandy-skeletal, siliceous, dysic, mesic Terric Haplosaprists high in organic deposits that are found in lake basins, back swamps, floodplains, and freshwater channels. The soils are classified in the dysic reaction class because pH in water dominantly ranges from 4.5 to 5.0. The zone from the soil surface to a depth of 200 cm is continuously saturated (endosaturation). The upper strata of Manahawkin Series from 0 to 76 cm is black broken face and rubbed muck; 10 percent fibers, less than 2 percent rubbed; moderate medium granular structure; mat of many fine roots; identifiable material is primarily herbaceous with a trace of woody fragments; 85-95 percent organic matter; extremely acid. Lower strata to 200 cm are gray sand, single grained and loose; strongly acid. (Source: NRCS Web Soil Survey.)Wild material2159118NA 88690
468NA 88691RJL2023NJ-004Ilex laevigata (Pursh) A. Gray New Jersey, United StatesNA2023COLLECTED11/10/2023Franklin Township: Piney Hollow Preservation Area (the site is owned by Franklin Township). This species was found below the dam at the far end of McCarthys Lakes. Plants accessed by foot along trails throughout property.39.57404200, -74.9252370028Plants growing in part-shade to sun of the moist to saturated areas below the McCarthys Lakes dam in the wetlands associated with the New Squankum Branch. The area appears to be inundated year-round with plants growing on low mounds surrounded by obligate wetland species. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Middle Miocene, Serravallian origin and characterized as an Cohansey Formation. It is comprised of sand, white to yellow with local gravel and clay. Locally stained red or orange brown by iron oxides and (or) cemented into large blocks of ironstone. Unweathered clay is typically dark gray, but commonly weathers white where interbedded with thin beds of ironstone. Unit is a complex of interfingering marine and nonmarine facies. Sand is typically medium grained and moderately sorted although it ranges from fine to very coarse grained and from poorly to well sorted. Sand consists of quartz and siliceous rock fragments. Some beds are locally micaceous. In general, the sand is crossbedded, although the style of crossbedding varies significantly with the paleoenvironment. Trough crossbedding predominates, especially in the nonmarine channel fill deposits, and the scale of the crossbeds varies from small to large. Maximum thickness in the map area is about 60 m; however, thickness is difficult to determine because of the irregular basal contact and extensive post-depositional erosion. There is as much as 18 m of relief along the basal contact. The basal contact is sharp, undulatory, and directly overlain by a thin gravel bed. The Cohansey Formation unconformably overlies the Kirkwood Formation and is found in channels cut down into the Kirkwood. Where the Kirkwood consists of sandy, light-colored sediments, the basal contact of the Cohansey is drawn below crossbedded sediments. Where the Kirkwood consists of dark-colored silty beds, the basal contact is drawn between light-colored Cohansey sediments and the underlying dark-colored sediments. The Cohansey was markedly thinned because of erosion prior to deposition of overlying units in the western and southern parts of the southern sheet. The unit has been extensively eroded and stripped from large areas of the New Jersey Coastal Plain, particularly in the central sheet where outliers are common. In spite of its widespread nature, the Cohansey is poorly exposed because of its loose sandy composition, which causes it to erode easily. (Source: USGS New Jersey Geologic Map Data.) The primary soil type in the collection area is classified as Manahawkin Series muck, 0 to 2 percent slopes, frequently flooded, Northern Coastal Plain. Manahawkin Series soils are very deep, very poorly drained soils that are sandy or sandy-skeletal, siliceous, dysic, mesic Terric Haplosaprists high in organic deposits that are found in lake basins, back swamps, floodplains, and freshwater channels. The soils are classified in the dysic reaction class because pH in water dominantly ranges from 4.5 to 5.0. The zone from the soil surface to a depth of 200 cm is continuously saturated (endosaturation). The upper strata of Manahawkin Series from 0 to 76 cm is black broken face and rubbed muck; 10 percent fibers, less than 2 percent rubbed; moderate medium granular structure; mat of many fine roots; identifiable material is primarily herbaceous with a trace of woody fragments; 85-95 percent organic matter; extremely acid. Lower strata to 200 cm are gray sand, single grained and loose; strongly acid. (Source: NRCS Web Soil Survey.)Wild material2159119NA 88691
469NA 88692RJL2023NJ-005Ilex opaca Aiton New Jersey, United StatesNA2023COLLECTED11/10/2023Franklin Township: Piney Hollow Preservation Area (the site is owned by Franklin Township). This species was found on as well as below the dam at the far end of McCarthys Lakes. Plants accessed by foot along trails throughout property.39.57423200, -74.9252560029Plants growing in part-shade to sun of the moist to saturated areas below the McCarthys Lakes dam in the wetlands associated with the New Squankum Branch. Plants were also found growing on the slopes of the dam in well-drained mesic to dry-mesic conditions. This species is clearly adaptable to a variety of conditions. Slope: 0 to 15 percent. Aspect: East. Elevation: 28 to 30 meters. NOTE: In addition to the soil type described below, this species was also growing on the slopes of a constructed earthen dam of unknown soil origin. The underlying geology of this area is of Middle Miocene, Serravallian origin and characterized as an Cohansey Formation. It is comprised of sand, white to yellow with local gravel and clay. Locally stained red or orange brown by iron oxides and (or) cemented into large blocks of ironstone. Unweathered clay is typically dark gray, but commonly weathers white where interbedded with thin beds of ironstone. Unit is a complex of interfingering marine and nonmarine facies. Sand is typically medium grained and moderately sorted although it ranges from fine to very coarse grained and from poorly to well sorted. Sand consists of quartz and siliceous rock fragments. Some beds are locally micaceous. In general, the sand is crossbedded, although the style of crossbedding varies significantly with the paleoenvironment. Trough crossbedding predominates, especially in the nonmarine channel fill deposits, and the scale of the crossbeds varies from small to large. Maximum thickness in the map area is about 60 m; however, thickness is difficult to determine because of the irregular basal contact and extensive post-depositional erosion. There is as much as 18 m of relief along the basal contact. The basal contact is sharp, undulatory, and directly overlain by a thin gravel bed. The Cohansey Formation unconformably overlies the Kirkwood Formation and is found in channels cut down into the Kirkwood. Where the Kirkwood consists of sandy, light-colored sediments, the basal contact of the Cohansey is drawn below crossbedded sediments. Where the Kirkwood consists of dark-colored silty beds, the basal contact is drawn between light-colored Cohansey sediments and the underlying dark-colored sediments. The Cohansey was markedly thinned because of erosion prior to deposition of overlying units in the western and southern parts of the southern sheet. The unit has been extensively eroded and stripped from large areas of the New Jersey Coastal Plain, particularly in the central sheet where outliers are common. In spite of its widespread nature, the Cohansey is poorly exposed because of its loose sandy composition, which causes it to erode easily. (Source: USGS New Jersey Geologic Map Data.) The primary soil type in the collection area is classified as Manahawkin Series muck, 0 to 2 percent slopes, frequently flooded, Northern Coastal Plain. Manahawkin Series soils are very deep, very poorly drained soils that are sandy or sandy-skeletal, siliceous, dysic, mesic Terric Haplosaprists high in organic deposits that are found in lake basins, back swamps, floodplains, and freshwater channels. The soils are classified in the dysic reaction class because pH in water dominantly ranges from 4.5 to 5.0. The zone from the soil surface to a depth of 200 cm is continuously saturated (endosaturation). The upper strata of Manahawkin Series from 0 to 76 cm is black broken face and rubbed muck; 10 percent fibers, less than 2 percent rubbed; moderate medium granular structure; mat of many fine roots; identifiable material is primarily herbaceous with a trace of woody fragments; 85-95 percent organic matter; extremely acid. Lower strata to 200 cm are gray sand, single grained and loose; strongly acid. (Source: NRCS Web Soil Survey.)Wild material2159120NA 88692
470NA 88697RJL2023NCSC-002Ilex amelanchier M. A. Curtis ex Chapm. North Carolina, United StatesNA2023COLLECTED11/12/2023Goose Pond Bay (Nature Conservancy-owned site): wetlands on the south side of McNeil Road approximately 0.64 kilometers west of junction with McIntosh Road.34.88254100, -79.0991570060Plants are growing in part-shade to sun of a Carolina Bay habitat with seasonally moist to wet, acidic conditions. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Cretaceous origin and characterized as Black Creek Formation. The formation is comprised of clay that is gray to black and lignitic. It contains thin beds and laminae of fine-grained micaceous sand and thick lenses of cross-bedded sand. This formation has glauconitic, fossiliferous clayey sand lenses in the upper part. (Source: USGS North Carolina Geologic Map Data.) The primary soil type in the collection area is classified as McColl loam soil. The McColl Series consists of poorly drained, slowly permeable soils that are shallow or moderately deep to a fragipan and very deep to bedrock. They formed in loamy sediments in oval depressions on the Coastal Plain. The soils are nearly level or concave. Runoff is frequently ponded. Slopes are less than 2 percent. These soils are strongly acid. (Source: NRCS Web Soil Survey.)Wild material2159125NA 88697
471NA 88698RJL2023NCSC-003Ilex laevigata (Pursh) A. Gray North Carolina, United StatesNA2023COLLECTED11/13/2023Sandhills Game Land: north side of Pulpwood Road approximately 0.11 km southwest of the junction with Tyner Road.34.95939500, -79.48676300114Plants growing in shade to part-shade of a spring fed wetland that is a mixed evergreen and deciduous woodland habitat. Slope: 3 to 10 percent. Aspect: Northeast. The underlying geology of this area is of Tertiary origin and characterized as Pinehurst Formation. It is comprised of sand that is medium- to coarse-grained, cross-bedding and rhythmic bands of clayey sand common, unconsolidated. (Source: USGS North Carolina Geologic Map Data.) The primary soil type in the area of this collection is classified as Pelion loamy sand, 6 to 10 percent slopes. Pelion Series soils are very deep, moderately well-drained soils with moderately slow to slow permeability. The upper 30 cm of the strata is sandy loam to sandy clay loam ranging from grayish brown to reddish yellow. This stratum has a weak fine granular structure and is very friable. It contains many fine roots and is moderately acid. Deeper strata to 2 m range from yellow to yellowish red sandy clay, often with masses of oxidized iron and few fine prominent light gray iron depletions; they are strongly acid. (Source: NRCS Web Soil Survey.)Wild material2159126NA 88698
472NA 88711RJL2023NCSC-016Ilex myrtifolia Walter South Carolina, United StatesNA2023COLLECTED11/14/2023Francis Marion National Forest: plants growing in wetlands/swamps on US Forest Service Road 223 approximately 0.57 kilometers southeast from the junction with Halfway Creek Road.33.06786200, -79.6776180011Plants are growing in shade to part-shade in the swamp in seasonally inundated acidic wetland habitat. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Pleistocene origin and characterized as an Socastee Formation. This is a low coastal formation in the Carolinas like Penholoway but younger and lower in altitude. It is comprised primarily of unconsolidated, fine-detrital clay. It is described as variegated quartzose sands, argillaceous sands, and clays, 5 m thick at the type section near Socastee Swamp. (Source: USGS South Carolina Geologic Map Data.) The primary soil type in this collection area is classified as Rutlege Series loamy fine sand. Rutlege Series soils are very deep, very poorly drained soils that are sandy, siliceous, thermic Typic Humaquepts. Upper strata soils (0-38 cm) are black loamy sand with weak medium granular structure. They are loose and very strongly acid. Lower strata (38-175 cm) range from dark gray to dark brown sand that is single grained, loose, and very strongly acid. (Source: NRCS Web Soil Survey.)Wild material2159139NA 88711
473NA 88713RJL2023NCSC-018Ilex opaca Aiton South Carolina, United StatesNA2023COLLECTED11/14/2023Francis Marion National Forest: plants growing in wetlands/swamps on US Forest Service Road 223 approximately 0.57 kilometers southeast from the junction with Halfway Creek Road.33.06783000, -79.6774910011Plants are growing in part shade to shade of a swamp in seasonally inundated acidic wetland habitat. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Pleistocene origin and characterized as an Socastee Formation. This is a low coastal formation in the Carolinas like Penholoway but younger and lower in altitude. It is comprised primarily of unconsolidated, fine-detrital clay. It is described as variegated quartzose sands, argillaceous sands, and clays, 5 m thick at the type section near Socastee Swamp. (Source: USGS South Carolina Geologic Map Data.) The primary soil type in this collection area is classified as Rutlege Series loamy fine sand. Rutlege Series soils are very deep, very poorly drained soils that are sandy, siliceous, thermic Typic Humaquepts. Upper strata soils (0-38 cm) are black loamy sand with weak medium granular structure. They are loose and very strongly acid. Lower strata (38-175 cm) range from dark gray to dark brown sand that is single grained, loose, and very strongly acid. (Source: NRCS Web Soil Survey.)Wild material2159141NA 88713
474NA 88716RJL2023NCSC-021Ilex myrtifolia Walter South Carolina, United StatesNA2023COLLECTED11/14/2023Francis Marion National Forest: plants growing in a pond cypress-water tupelo swamp on the west side of Thompson Branch Road just south of the junction with Round Pond Road B.33.15167800, -79.6025370031Plants are growing in part-shade to shade as understory trees among the pond cypress-water tupelo forest throughout the acid swamp which is seasonally inundated. The overstory is largely deciduous and comprised of pond cypress and water tupelo, while the understory is a mix of evergreen and deciduous broadleaved shrubs and understory trees. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Pleistocene origin and characterized as a Socastee Formation. This is a low coastal formation in the Carolinas like Penholoway but younger and lower in altitude. It is comprised primarily of unconsolidated, fine-detrital clay. It is described as variegated quartzose sands, argillaceous sands, and clays, 5 m thick at the type section near Socastee Swamp. (Source: USGS South Carolina Geologic Map Data.) The primary soil type in this collection area is classified as Pickney Series loamy fine sand. The Pickney Series consists of very deep, very poorly drained, persistently wet, permeable soils that formed from marine or fluvial sediments. Slopes range from 0 to 2 percent. The upper strata soil, 0-75 cm, is black loamy fine sand that is very friable and extremely acid. The lower strata, 75-120 cm is dark gray fine sand with occasional white mottling, loose, and moderately acid. (Source: NRCS Web Soil Survey.)Wild material2159144NA 88716
475NA 88717RJL2023NCSC-022Ilex opaca Aiton South Carolina, United StatesNA2023COLLECTED11/14/2023Francis Marion National Forest: plants growing in a pond cypress-water tupelo swamp on the west side of Thompson Branch Road just south of the junction with Round Pond Road B.33.15161300, -79.6021790022Plants are growing in part-shade to shade as understory trees among the pond cypress-water tupelo forest, mostly along the edges of the acid swamp which is seasonally inundated. The overstory is largely deciduous and comprised of pond cypress and water tupelo, while the understory is a mix of evergreen and deciduous broadleaved shrubs and understory trees. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Pleistocene origin and characterized as a Socastee Formation. This is a low coastal formation in the Carolinas like Penholoway but younger and lower in altitude. It is comprised primarily of unconsolidated, fine-detrital clay. It is described as variegated quartzose sands, argillaceous sands, and clays, 5 m thick at the type section near Socastee Swamp. (Source: USGS South Carolina Geologic Map Data.) The primary soil type in this collection area is classified as Pickney Series loamy fine sand. The Pickney Series consists of very deep, very poorly drained, persistently wet, permeable soils that formed from marine or fluvial sediments. Slopes range from 0 to 2 percent. The upper strata soil, 0-75 cm, is black loamy fine sand that is very friable and extremely acid. The lower strata, 75-120 cm is dark gray fine sand with occasional white mottling, loose, and moderately acid. (Source: NRCS Web Soil Survey.)Wild material2159145NA 88717
476NA 88718RJL2023NCSC-023Ilex decidua Walter South Carolina, United StatesNA2023COLLECTED11/15/2023Waccamaw National Wildlife Refuge: U.S. Highway 701, Yauhannah Landing on the east side of the Great Pee Dee River. Plants growing in low woods on the north side of the landing.33.66208900, -79.153050002Plants growing in shade to part-shade of a moist to wet woodland habitat that is mostly deciduous and evergreen. Portions of the woodland have depressions with year-round standing water. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Quaternary origin and characterized as Alluvial Valley Swamp. This formation is unconformable on all underlying units. It is composed of fluvial sand and gravel at base, grading upwards into fine sands and silts, and local occasional peat. May be overrun with recent sediments from forest cutting and agriculture. (Source: USGS South Carolina Geologic Map Data.) The primary soil type in this collection area is classified as Johnston Series loam. The Johnson Series consists of very deep, very poorly drained coarse-loamy, siliceous, active, acid, thermic Cumulic Humaquepts. The upper 0-75 cm is black, mucky loam that is friable and very strongly acid. The lower stratum to 1.75 m is dark gray, loamy fine sand, single grained, loose and very strongly acid. (Source: NRCS Web Soil Survey.)Wild material2159146NA 88718
477NA 88719RJL2023NCSC-024Ilex laevigata (Pursh) A. Gray South Carolina, United StatesNA2023COLLECTED11/15/2023Little Pee Dee Heritage Preserve/Wildlife Management Area (South Carolina Department of Natural Resources Public Lands, on South Carolina Highway 917).34.14123300, -79.1953210013Plants growing in high understory shade to part-shade of a moist to relatively dry pine forest with occasional deciduous trees. The litter layer in the woodlands is very deep with longleaf pine needles, while the edges of the woodlands are much drier with exposed white sand and little to no litter layer. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Pleistocene origin and characterized as Waccamaw Formation. This formation is another Carolina coastalized terrace that is deeply weathered. It is characterized by unconsolidated, coarse-detrital, soft sand limestones and loose gray to buff fine quartz sands in which occasional small quartz pebbles are present. It also includes sedimentary, carbonate, soft limestones. (Source: USGS South Carolina Geologic Map Data.) The primary soil type in this collection area is classified as Lynn Haven Series sand. The Lynn Haven Series consists of very deep, poorly and very poorly drained, moderate or moderately rapid, permeable soils in low areas and depressions in Atlantic Flatwoods and the Gulf. They formed in thick deposits of sandy marine sediments. Th Lynn Haven Series soil is approximately 2 m thick and ranges from black near the surface to yellow brown in the deepest strata. Most of the strata are comprised of fine sands that are friable with the deepest strata also including a small percentage of organic matter. Reaction ranges from extremely acid to strongly acid throughout the profile. The water table is at 0-15 cm for periods of 2 to 6 months annually and within a depth 1 m for more than 6 months during most years; during extended dry periods it is below 1 meter. Depressional areas are ponded for long duration in most years. (Source: NRCS Web Soil Survey.)Wild material2159147NA 88719
478NA 88721RJL2023NCSC-026Ilex opaca Aiton South Carolina, United StatesNA2023COLLECTED11/15/2023Little Pee Dee Heritage Preserve/Wildlife Management Area (South Carolina Department of Natural Resources Public Lands, on South Carolina Highway 917).34.14091500, -79.1955520013Plants growing in high understory shade to part-shade of a moist to relatively dry pine forest with occasional deciduous trees. The litter layer in the woodlands is very deep with longleaf pine needles, while the edges of the woodlands are much drier with exposed white sand and little to no litter layer. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Pleistocene origin and characterized as Waccamaw Formation. This formation is another Carolina coastalized terrace that is deeply weathered. It is characterized by unconsolidated, coarse-detrital, soft sand limestones and loose gray to buff fine quartz sands in which occasional small quartz pebbles are present. It also includes sedimentary, carbonate, soft limestones. (Source: USGS South Carolina Geologic Map Data.) The primary soil type in this collection area is classified as Lynn Haven Series sand. The Lynn Haven Series consists of very deep, poorly and very poorly drained, moderate or moderately rapid, permeable soils in low areas and depressions in Atlantic Flatwoods and the Gulf. They formed in thick deposits of sandy marine sediments. Th Lynn Haven Series soil is approximately 2 m thick and ranges from black near the surface to yellow brown in the deepest strata. Most of the strata are comprised of fine sands that are friable with the deepest strata also including a small percentage of organic matter. Reaction ranges from extremely acid to strongly acid throughout the profile. The water table is at 0-15 cm for periods of 2 to 6 months annually and within a depth 1 m for more than 6 months during most years; during extended dry periods it is below 1 meter. Depressional areas are ponded for long duration in most years. (Source: NRCS Web Soil Survey.)Wild material2159149NA 88721
479NA 88723RJL2023NCSC-028Ilex myrtifolia Walter North Carolina, United StatesNA2023COLLECTED11/15/2023Juniper Creek State Game Lands (along Brunswick County Road 1340): swamps/wetlands associated with Juniper Creek.34.12438700, -78.3931070013Plants growing in shade to part-shade of a swamp habitat along Juniper Creek. At the time of collection, the swamp was relatively dry with water found only in deep depressions. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Cretaceous origin and characterized as Peedee Formation - Black Creek Group, undivided. The Peedee Formation is primarily unconsolidated, coarse-detrital, sand that is dark-green or gray, finely micaceous, more or less glauconitic, and argillitic sand, many layers of which are calcareous (impure limestone). Irregular concretionary masses of impure calcium carbonate occur in places. Dark marine clays are interstratified with sand. The Black Creek Group consists of irregularly bedded, laminated, carbonaceous clays and thin laminae and lenses of sand; lignite and pyrite present; glauconitic in places; massive interbedded layers of glauconitic sand present toward top of unit. Light-colored clays and coarse arkosic sands also found. (Source: USGS North Carolina Geologic Map Data.) The primary soil type in this collection area is classified as Dorovan Series muck. The Dorovan Series consists of very deep, very poorly drained, moderately permeable soils on densely forested flood plains, hardwood swamps, and depressions in the Atlantic Coast Flatwoods, Eastern Gulf Coast Flatwoods, and Southern Coastal Plain Major Land Resource Areas. They formed in highly decomposed acid-organic materials and have slopes that are less than 1 percent. (Source: NRCS Web Soil Survey.)Wild material2159151NA 88723
480NA 88725RJL2023NCSC-030Ilex myrtifolia Walter North Carolina, United StatesNA2023COLLECTED11/15/2023Boiling Spring Lakes, NC: Plants growing along unused road of potential new development on Baden Street, between Camden Street and Midwood Street. The site is accessed from North Carolina Highway 87 (George II Highway SE) by turning west onto Brunswick Road and right onto Baden Street at the first intersection.34.01935700, -78.0611050016Plants growing in part-shade along the road, edges of ditches and managed edges of longleaf pine forest in well-drained, but seasonally moist habitat with very little organic litter layer. It seems that low competition and exposed sandy conditions allows this plant to thrive. Slope: N/A. Aspect: N/A. The underlying geology of this area is of Pleistocene origin and characterized as Waccamaw Formation. This formation is another Carolina coastalized terrace that is deeply weathered. It is characterized by unconsolidated, coarse-detrital, soft sand limestones and loose gray to buff fine quartz sands in which occasional small quartz pebbles are present. It also includes sedimentary, carbonate, soft limestones. (Source: USGS North Carolina Geologic Map Data.) The primary soil type in this collection area is classified as Leon Series fine sand. The Leon series consists of very deep, very poorly and poorly drained, moderately rapid to moderately slowly permeable soils on upland flats, depressions, stream terraces and tidal areas. They formed in sandy marine sediments of the Eastern Gulf Coast Flatwoods, the Atlantic Coast Flatwoods and to a lesser extent in the Southern Coastal Plain. The upper horizon (0-75 cm) is comprised of a high percent of blac and lower percentage of light gray sand that is very friable and very strongly acid. The lower horizon (75-270 cm) has a range of gray, dark brown and yellowish-brown sands that range from fine to coarse and are also very strongly acid. (Source: NRCS Web Soil Survey.)Wild material2159153NA 88725
481NA 88686TJK2023FL-033Ilex glabra (L.) A. Gray Florida, United StatesNA2023COLLECTED11/18/2023Navarre, Florida: Adams St. cul-de-sac, on nine-acre private property. Infrastructure development began and was terminated due to the lack of any engineering or environmental site planning (designated wetland status). It has remained idle for a number of years.30.40970000, -86.9298000011Wet pine flatwood transitioning to a shrub bog habitat under fire suppression. Slope: 3.5%. Aspect: West. Elevation: 36 feet.Wild material2159098NA 88686
482NA 88688TJK2023FL-035Ilex decidua Walter Florida, United StatesNA2023COLLECTED11/19/2023Eglin Air Force Base, Choctaw West; Range Rd 813, approximately 0.3 miles north of Range Rd 213 - Choctaw Nolf Airport Rd. Growing in flat west of Weaver Creek.30.51790000, -86.9323000034Damp (not wet) flat west of creek. Slope: less than 2%. Aspect: West. Elevation: 113 feet.Wild material2159100NA 88688
483NA 87576'Golden Nellie'Ilex hybr.NANot Available2023CultivarRecorded at the U.S. National Arboretum as Ilex 'Golden Nellie'. Received as Ilex 'Whoa Nellie'. Rooted cutting. Holly Society of America test holly for 2024.2165744NA 87576
484NA 88858TRx231112-01MDIlex opaca Aiton Maryland, United StatesNA2023COLLECTED11/12/202338.98349100, -76.9171440022Wild material2159075NA 88858
485NA 88681RJL2023PA-015Ilex verticillata (L.) A. Gray Pennsylvania, United StatesNA2023COLLECTED10/30/2023Wetlands near Phillips Creek: seed collected from plants on the north and south (further east) side of Pennsylvania Route 188 approximately 6.9 kilometers east of the junction with Pennsylvania Route 487.41.30353100, -76.22452800381Boggy to moist habitats associated with wetlands of Phillips Creek in shade to full sun next to the road. Slope: 0%. Aspect: various. The underlying geology of this area is of Devonian origin and characterized as the Catskill Formation. It is comprised of grayish-red sandstone, siltstone, shale, and mudstone; units of gray sandstone occur in upper part; lithologies in upper part arranged in fining-upward cycles. In the Altoona area, the Catskill Formation is mapped as the Duncannon, Sherman Creek, and Irish Valley Members, which are described under "Central and Eastern Pennsylvania" provinces. (Source: USGS Pennsylvania Geologic Map Data). The primary soil type in the collection area is classified as Wellsboro Series. The Wellsboro Series consists of very deep, moderately well- and somewhat poorly-drained soils formed in till derived from red sandstone, siltstone, and shale. Slope ranges from 0 to 50 percent. Saturated hydraulic conductivity is moderately high or high in the mineral surface layer, subsurface layer, and upper part of the subsoil; and low or moderately low in the lower part of the subsoil (fragipan) and the substratum. Solum thickness is greater than 102 cm. Depth to the fragipan ranges from 36 to 76 cm. Depth to bedrock is 152 cm or more. Rock fragments of subangular and rounded sandstone, siltstone or shale range from 5 to 40 percent in the A and B horizons, and from 15 to 50 percent in the Bx and C horizons. Typically, rock fragments average about 5 to 25 percent by volume above the fragipan and 15 to 40 percent by volume in and below the fragipan. Reaction commonly ranges from very strongly acid through moderately acid. (Source: NRCS Web Soil Survey).Wild material2159072NA 88681
486NA 88669TXWL2023-012Ilex decidua Walter Texas, United StatesNA2023COLLECTED10/21/2023County Road 3218 en route to Camp Wohaleta, north of Caddo National Grasslands.33.75626700, -95.97888900Roadside.Wild material2159044NA 88669
487NA 88665RJL2023PA-002Ilex verticillata (L.) A. Gray Pennsylvania, United StatesNA2023COLLECTED10/01/2023Woodlands along Pine Creek near the Harmonyville Road bridge.40.18764500, -75.76732200141Moist to mucky, wet woodland conditions along Pine Creek. Much of the site is very wet due to recent rains; though, earlier in the year, the soils were only moist. Plants grow in both well-drained mesic conditions as well as seasonally wet conditions. They do not grow in standing water anywhere in this location. Plants in this population are found mostly in the shade and partial shade of deciduous trees; there are occasional gaps that offer higher light conditions as well. Slope: 0-2%. Aspect: N/A. The underlying geology of this area is of Precambrian origin and characterized as the Graphitic Felsic Gneiss. This includes Pickering Gneiss and small areas of marble; dominantly quartz and feldspar with varying amounts of graphite and various metamorphic minerals; medium grained, light to dark gray and greenish gray; sedimentary origin. (Source: USGS Pennsylvania Geologic Map Data). The primary soil type in the area of this collection is classified as Hatboro Silt Loam Series. The Hatboro Series consists of very deep and poorly drained soils formed in alluvium derived from metamorphic and crystalline rock. They are on flood plains. Slopes range from 0 to 3 percent. Saturated hydraulic conductivity is moderately high to high. Solum thickness ranges from 50 to 150 cm. Depth to bedrock ranges from 1.5 to 3 meters or more. Organic carbon decreases irregularly with depth or is greater than 0.2 percent directly above any strongly contrasting C horizon. The depth to strongly contrasting sand and gravel is more than 100 cm or the transition is greater than 12.5 cm. Content of gravel ranges from 0 to 10 percent in the solum and from 0 to 80 percent in the C horizon. Mica flakes are common in the solum, and the C horizon contains thin lenses of mica. Reaction ranges from very strongly acid through neutral to a depth of 75 cm and from moderately acid through slightly acid below 75 cm. (Source: NRCS Web Soil Survey.)Wild material2158814NA 88665
488NA 88658TJK2023FL-012Ilex verticillata (L.) A. Gray Florida, United StatesNA2023COLLECTED09/10/2023Lower Yellow River.30.62868600, -86.807480003Beautiful riverine system. Elevation: 13 feet. Slope: 0 (none). Aspect: Northwest.Wild material2158792NA 88658
489NA 88659TJK2023FL-013Ilex sp.Florida, United StatesNA2023COLLECTED09/15/2023Lower Yellow River.30.58170000, -86.917700001Elevation: 3 feet. Slope: 0 percent. Aspect: North.Wild material2158793NA 88659
490NA 88606KY23-008Ilex montana Torr. & A. Gray Kentucky, United StatesNA2023COLLECTED09/09/2023Black Mountain summit (second location, bulked with the first: Black Mountain; lat/long 36. 91614, -82. 89873)36.91434000, -82.894630001261Moist woods. Elevation: 4138 feet. Aspect: West. (Second location, bulked with first: Elevation: 4071 feet. Aspect: West.)Wild material2158698NA 88606
491NA 88621KY23-025aIlex ambigua Torr. Kentucky, United StatesNA2023COLLECTED09/10/2023The Gulf - Natural Arch Scenic Area Panoramic Trail.36.84417000, -84.52833000395Ridgeline. Elevation: 1295 feet. Aspect: South.Wild material2158713NA 88621
492NA 88622KY23-025bIlex ambigua Torr. Kentucky, United StatesNA2023COLLECTED09/10/2023The Gulf - Natural Arch Scenic Area Panoramic Trail.36.84417000, -84.52833000395Ridgeline. Elevation: 1295 feet. Aspect: South.Wild material2158714NA 88622
493NA 88640KY23-043Ilex verticillata (L.) A. Gray Kentucky, United StatesNA2023COLLECTED09/11/2023Daniel Boone National Forest: Bee Rock Loop Trail along Rockcastle River.37.04179000, -84.31805000233Riverscour grassland. Elevation: 765 feet. Aspect: East.Wild material2158732NA 88640
494NA 87338NA 87338Ilex purpurea Hassk. NANot Available2023Uncertain improvement statusFull name as recorded at U.S. National Arboretum: Ilex purpurea. Seedling (at least several years old) that appeared just north of the Kingman Lake Overlook loop guard rail amid invasive vines and shrubs, near some old Ilex opaca. Probably deposited from seed of NA 42133 from the other side of the collection, or maybe less likely from NA 69765 on the Anacostia River Flood Plain, if it is able to produce viable seed (no males are nearby). Since we have few plants of this taxon and parents are fairly likely to be of wild origin, this might be worth keeping and observing further. -SBL2155331NA 87338
495NA 76317'William Hawkins'Ilex opaca Aiton NANot Available2023CultivarFull name as recorded at U.S. National Arboretum: Ilex opaca 'William Hawkins'.2150665NA 76317
496NA 76027(undetermined cultivar)Ilex crenata Thunb. NANot Available2023Uncertain improvement statusFull name as recorded at U.S. National Arboretum: Ilex crenata (undetermined cultivar).2150578NA 76027
497NA 74887NA 74887Ilex ×koehneana Loes. NANot Available2023Uncertain improvement statusFull name as recorded at U.S. National Arboretum: Ilex x koehneana. Looks like latifolia. [Actually, it appears to be I. x koehneana, but does not quite match any cultivars in our collections as of 2025; it could be an unnamed seedling selection, or a cultivar otherwise absent from USNA at this time.]2150193NA 74887
498NA 73322NA 73322Ilex opaca Aiton NANot Available2023Uncertain improvement statusFull name as recorded at U.S. National Arboretum: Ilex opaca. Old USNA plant. Form received: unknown propagule type.2149598NA 73322
499NA 73320NA 73320Ilex sp.NANot Available2023Uncertain improvement statusFull name as recorded at U.S. National Arboretum: Ilex sp.. Old USNA plant. Form received: unknown propagule type.2149596NA 73320