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Taxon: Cucumis sativus L. var. hardwickii (Royle) Alef.

Genus: Cucumis
Subgenus: Cucumis
Section: Cucumis
Family: Cucurbitaceae
Tribe: Benincaseae
Nomen number: 104944
Place of publication: Landw. Fl. 196. 1866
Link to protologue:
  • • Alefeld's combination is valid under the varietal rank
  • • the introduction (page V) of the original publication explains the application of the varietal rank
Name Verified on: 06-Feb-1996 by ARS Systematic Botanists.
Accessions: 9 (7 active, 6 available) in National Plant Germplasm System (Map)

Autonyms (not in current use), synonyms and invalid designations:

(≡ homotypic synonym, = heterotypic synonym, - autonym, I invalid designation)

Common names:

  • xi nan ye huang gua  (Source: F ChinaEng) - Transcribed Chinese

Distributional Range:


    • CHINA: China [Guizhou Sheng, Yunnan Sheng, Guangxi Zhuangzu Zizhiqu]

    • INDIAN SUBCONTINENT: India (n.), Nepal
    • INDO-CHINA: Myanmar (n.), Thailand (n.)


  1. Chinese Academy of Sciences. 1959-. Flora reipublicae popularis sinicae.
  2. De Wilde, W. J. J. O. & B. E. E. Duyfjes. 2007. The wild species of Cucumis L. (Cucurbitaceae) in South-East Asia. Adansonia sér. 3, 29:246-247.
  3. Garcia-Mas, J. et al. 2004. Phylogenetic relationships among Cucumis species based on the ribosomal internal transcribed spacer sequence and microsatellite markers. Pl. Syst. Evol. 248:191-203.
  4. Hara, H. et al. 1978-1982. An enumeration of the flowering plants of Nepal. Note: lists as Cucumis sativus var. hardwickii (Royle) Kitam.
  5. Jeffrey, C. 1980. Further notes on Cucurbitaceae: V. The Cucurbitaceae of the Indian Subcontinent. Kew Bull. 34:802. Note: lists as Cucumis sativus var. hardwickii (Royle) Gabaev.
  6. Kho, Y. O. et al. 1980. Interspecific hybridization in Cucumis L. II. The crossability of species, and investigation of in vivo pollen tube growth and seed set. Euphytica 29:661-672. Note: this study examined USDA accession PI 215589
  7. Kirkbride, J. H., Jr. 1993. Biosystematic monograph of the genus Cucumis (Cucurbitaceae) 119. Note: = Cucumis sativus L.
  8. Lebeda, A. et al. 2007. Chapter 8. Cucurbits (Cucurbitaceae; Cucumis spp., Cucurbita spp., Citrullus spp.). Genetic resources, chromosome engineering, and crop improvement 3:271-376. Note: this review recognized var. hardwickii as part of the primary gene pool of the cultivated cucumber
  9. Meglic, V. et al. 1996. Genetic diversity in cucumber (Cucumis sativus L.): I. A reevaluation of te U.S. germplasm collection. Genet. Resources Crop Evol. 43:533-546.
  10. Porcher, M. H. et al. Searchable World Wide Web Multilingual Multiscript Plant Name Database (MMPND) (on-line resource).
  11. Renner, S. S. 2017. A valid name for the Xishuangbanna gourd, a cucumber with carotene-rich fruits. PhytoKeys 85:87-94. Note: this variety is recognized as source of the progenitor population of cucumber
  12. Sebastian, P. et al. 2010. Cucumber (Cucumis sativus) and melon (C. melo) have numerous wild relatives in Asia and Australia, and the sister species of melon is from Australia. Proc. Natl. Acad. Sci. U.S.A. 107:14269-14273. Note: lists as Cucumis sativus var. hardwickii (Royle) Gabaev.
  13. Smitinand, T. & K. Larsen, eds. 1970-. Flora of Thailand.
  14. Wu Zheng-yi & P. H. Raven et al., eds. 1994-. Flora of China (English edition).
  15. Yang, L. et al. 2012. Chromosome rearrengements during domestication of cucumber as revealed from high-density genetic mapping and draft genome assembly. Plant J. 71:895-906.
  16. Zhao, X. et al. 2011. Comparison of the distribution of the repetitive DNA sequences in three variants of Cucumis sativus reveals their phylogenetic relationships. J. Genet. Genomics 38:39-45. Note: formerly Acta Genetica Sinica

Check other web resources for Cucumis sativus L. var. hardwickii (Royle) Alef. :

  • Flora of China: Online version from Harvard University
  • TROPICOS: Nomenclatural and Specimen Database of the Missouri Botanical Garden
  • ePIC: Electronic Plant Information Centre of Royal Botanic Gardens, Kew
  • AGRICOLA: Article Citation Database or NAL Catalog of USDA's National Agricultural Library
  • Entrez: NCBI's search engine for PubMed citations, GenBank sequences, etc.
  • PubAg: USDA's National Agricultural Library database of full-text journal articles and citations on the agricultural sciences.

Cite as: USDA, Agricultural Research Service, National Plant Germplasm System. 2020. Germplasm Resources Information Network (GRIN-Taxonomy).
National Germplasm Resources Laboratory, Beltsville, Maryland. URL: https://npgsweb.ars-grin.gov/gringlobal/taxonomydetail.aspx?id=104944. Accessed 5 July 2020.